[Crustacea • 2017] A Case of Appalachian Endemism: Revision of the Cambarus robustus complex (Decapoda: Cambaridae) in the Kentucky and Licking River Basins of Kentucky, USA, with the Description of Three New Species ---ScRaBBlE

Cambarus (Puncticambarus) guenteri

 Loughman, Henkanaththegedara, Fetzner & Thoma, 2017

  

 DOI: 10.11646/zootaxa.4269.4.4 

Abstract

The amazing levels of freshwater biodiversity found in the Appalachian Mountains of the eastern United States are among the highest recorded globally. Localized endemics make up much of this diversity, with numerous fish, freshwater mussels, salamanders and crayfish often being restricted to a single watershed, and in some instances, subwatersheds. Much of this diversity is the product of the processes of vicariance and historical stream drainage patterns. Herein, we describe three new crayfish species, all previously members of the Cambarus robustus complex, which occur in the Appalachian portion of the Kentucky and Licking river basins in Kentucky, USA. All three species differ from each other morphologically, genetically, and zoogeographically, fulfilling the requirements of the integrated species concept. Cambarus guenteri occurs in the southern tributaries of the Kentucky River mainstem as well as throughout the South Fork Kentucky River. Cambarus taylori is a narrow endemic, which only occurs in the Middle Fork Kentucky River. Cambarus hazardi, which has the widest distribution of the three new species, occurs in the North Fork Kentucky River, Red River, and upper reaches of the Licking River basin. Stream piracy events between the Cumberland and South Fork Kentucky River, as well as the Licking, Red and North Fork Kentucky rivers, are theorized to be important in the evolution of this complex. Cambarus guenteri is proposed as currently stable, though both C. taylori and C. hazardi are considered imperiled at this time due to habitat destruction throughout both of their respective ranges.

Keywords: Cambarus, endemicity, Kentucky, new species, systematics, taxonomy, Crustacea

Cambarus (Puncticambarus) guenteri, new species

Etymology. It is with great pleasure that we name this species in honor of Dr. Guenter A. Schuster, Professor Emeritus, Eastern Kentucky University. Dr. Schuster dedicated his professional career to teaching undergraduates, graduate students and professional biologists about the diversity, biology, and conservation of freshwater invertebrates, with a particular influence on biologists from the bluegrass state, and he is a coauthor of the definitive work on the crayfishes of Kentucky. He instilled in those he educated a sense of wonder and appreciation for these organisms, which has undoubtedly led to the conservation of more than one freshwater mussel or crayfish. The common name for C. guenteri is the Redbird Crayfish in reference to the Redbird River drainage where large populations of C. guenteri occur.

Common name. Redbird Crayfish.

Cambarus (Puncticambarus) hazardi, new species

Etymology. Cambarus hazardi is named after Commodore Oliver Hazard Perry, noted American Naval Commander, and the name sake of Perry County Kentucky and the County Seat, Hazard Kentucky. The common name Brawny Crayfish is in reference to the powerfully built stature of C. hazardi.

Common name. Brawny Crayfish

Cambarus (Puncticambarus) taylori, new species

Etymology. It is our honor and privilege to name this crayfish after Dr. Christopher A. Taylor from the Illinois Natural History Survey. Dr. Taylor has been one of the most active crayfish researchers in the United States for the past two decades and a leader in crayfish conservation, co-authored the seminal work on Kentucky’s crayfishes, Crayfishes of Kentucky, and has been instrumental in bringing the conservation concerns of North America’s crayfishes to light with his many publications. The common name Cutshin Crayfish is in reference to Cutshin Creek watershed, which harbors the species.

Common name. Cutshin Crayfish

Zachary J. Loughman, Sujan M. Henkanaththegedara, James W. Fetzner, Jr. and Roger F. Thoma. 2017. A Case of Appalachian Endemism: Revision of the Cambarus robustus complex (Decapoda: Cambaridae) in the Kentucky and Licking River Basins of Kentucky, USA, with the Description of Three New Species.  Zootaxa. 4269(4); 460-494.  DOI: 10.11646/zootaxa.4269.4.4

ResearchGate.net/publication/317098813_A_case_of_Appalachian_endemism_Revision_of_the_Cambarus_robustus_complex_Decapoda_Cambaridae_in_the_Kentucky_and_Licking_River_basins_of_Kentucky_USA_with_the_description_of_three_new_species

Carnegie researcher discovers three more species of Appalachian crayfish  post-gazette.com/news/science/2017/08/05/Carnegie-museum-researcher-discover-species-Appalachian-crayfish-crawdads-crawfish-Zootaxa-James-Fetzner-Jr/stories/201708050034 via @PittsburghPG

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

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شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Crustacea • 2017] Procambarus virginalis • The Marbled Crayfish (Decapoda: Cambaridae) represents An Independent New Species ---ScRaBBlE

Procambarus virginalis  Lyko, 2017

 DOI: 10.11646/zootaxa.4363.4.6 

Abstract

Marbled crayfish are a globally expanding population of parthenogenetically reproducing freshwater decapods. They are closely related to the sexually reproducing slough crayfish, Procambarus fallax, which is native to the southeastern United States. Previous studies have shown that marbled crayfish are morphologically very similar to P. fallax. However, different fitness traits, reproductive incompatibility and substantial genetic differences suggest that the marbled crayfish should be considered an independent species. This article provides its formal description and scientific name, Procambarus virginalis sp. nov.

Keywords: Crustacea, parthenogenesis, annulus ventralis, genetic analysis, mitochondrial DNA

FIGURE 2. Procambarus virginalis new species holotype, dorsal views.  

Procambarus virginalis sp. nov.

Diagnosis. Body pigmented, eyes well developed. Rostrum with marginal spine and lacking median carina. Carapace with cervical spine. Areola 5 to 7 times as long as wide, constituting 30 to 35 percent carapace length (Tab. 1). Suborbital angle obtuse and weak. Postorbital ridge well developed with cephalic spine. Hepatic area punctate. Antennal scale approximately 2.5 times as long as wide, widest at midlength. Annulus ventralis bellshaped, about 1.6 times as broad as long, bisected by narrow furrow leading caudally into median depression. Sinus originating on median line, continuing longitudinally on anterior half, then curving dextrally before curving caudally before continuing and terminating on median line. Sternum immediately cephalic to annulus with no tubercles or projections and not overhanging annulus. Unadorned bell-shaped postannular sclerite with central longitudinal furrow, width similar to annulus. First pleopods present. Mitochondrial DNA with guanine and cytosine at positions 8754 and 8783, respectively (GenBank accession number KT074364, see Tab. 2 for a list of discriminatory genetic variants).

Etymology. The name Procambarus virginalis is derived from the preliminary designation Procambarus fallax forma virginalis (Martin et al., 2010) and reflects the species' unique parthenogenetic mode of reproduction. 

Remarks. The oldest known record of P. virginalis is from a German biologist and hobby aquarist. In a personal conversation with the author (February 2017), he recalled obtaining an uncharacterized batch of "Texas crayfish" from a pet trader specializing in American insects and other invertebrates, at a trade fair in Frankfurt (Germany) in 1995. Animal numbers of this original stock increased rapidly and animals were subsequently distributed to other German aquarists, eventually reaching commercial traders and pet stores. The absence of male animals and the parthenogenetic mode of reproduction were soon recognized among aquarists and subsequently confirmed in the first scientific description of marbled crayfish (Scholtz et al., 2003).

Frank Lyko. 2017. The Marbled Crayfish (Decapoda: Cambaridae) represents An Independent New Species. Zootaxa. 4363(4); 544–552. DOI: 10.11646/zootaxa.4363.4.6

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

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شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Crustacea • 2017] Cambarus (C.) appalachiensis • A New Species of Crayfish (Decapoda: Cambaridae) from the New River Basin of Virginia and West Virginia, USA ---ScRaBBlE

Cambarus (Cambarus) appalachiensis

Loughman, Welsh & Thoma, 2017

  DOI: 10.11646/zootaxa.4243.3.2 

Abstract

Cambarus (Cambarus) appalachiensis is a stream-dwelling crayfish endemic to the greater New River basins of Virginia and West Virginia. The new species is morphologically most similar to Cambarus sciotensis. Cambarus appalachiensis can be differentiated from C. sciotensis by its more elongated chelae which possess a single mesial row of tubercles, reduced to no tuberculation on the dorsal-longitudinal ridge of the dactyl, and reduced lateral impression. Cambarus sciotensis has a more subrectangular chelae with two rows of mesial margin tubercles on the chelae, as well as both a pronounced dorsal-longitudinal ridge and pronounced lateral impression. Several chelae meristic ratios also differentiate C. appalachiensis from C. sciotensis. Within the New, Gauley, and lower portions of the Greenbrier basins C. appalachiensis is the dominant tertiary burrowing Cambarus species, and as such, is considered stable across its range.

Keywords: Crustacea, Crayfish; New River; Appalachian Mountains

 Zachary J. Loughman, Stuart A. Welsh and Roger F. Thoma. 2017.
 Cambarus (C.) appalachiensis, A New Species of Crayfish (Decapoda: Cambaridae) from the New River Basin of Virginia and West Virginia, USA.
 Zootaxa. 4243(3); 432-454. DOI: 10.11646/zootaxa.4243.3.2

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Crustacea • 2019] Lacunicambarus chimera • A New Species of Burrowing Crayfish (Decapoda: Cambaridae) from Illinois, Indiana, Kentucky, and Tennessee ---ScRaBBlE

Lacunicambarus chimera Glon & Thoma,

in Glon, Thoma, Daly & Freudenstein, 2019. 

Crawzilla Crawdad  ||  DOI:  10.11646/zootaxa.4544.4.1

Photo by Guenter Schuster.

Abstract

Lacunicambarus diogenes (Girard 1852) was, until recently, considered to be one of the most widely distributed North American crayfish species, occurring in 31 U.S. States and one Canadian province east of the North American Rocky Mountains. Glon et al. (2018) investigated this claim and found that L. diogenes sensu lato was actually a species complex. The authors redescribed L. diogenes and restricted its range to the Atlantic Coastal Plain and Piedmont ecoregions of eastern North America. In doing so, they also revealed the existence of several probable undescribed species of Lacunicambarus that were previously considered to be L. diogenes. Here, we use morphological and molecular techniques to distinguish and describe one of these species: Lacunicambarus chimera sp. nov., a large primary burrowing crayfish found in parts of the Lower Mississippi, Ohio, Tennessee and Upper Mississippi River Basins. Lacunicambarus chimera is morphologically similar to L. diogenes, from which it can be distinguished by the greater number of spines on the ventrolateral margin of its merus, its wider antennal scale terminating in a short spine, and the presence of a single longitudinal stripe on the dorsal side of its abdomen. We also provide an updated key to Lacunicambarus.

Keywords: Crustacea, burrowing crayfish, systematics, taxonomy, revision, North America, freshwater, Old Ohio River

FIGURE 6. Dorsal view of Form I holotypic male of Lacunicambarus chimera (OSUMC 10650).

Photo by Guenter Schuster.

Taxonomy 

Family Cambaridae Hobbs 1942 

Genus Lacunicambarus (Hobbs 1969) 

Lacunicambarus chimera Glon & Thoma sp. nov.

Cambarus obesus Forbes 1876:6 [in part]. 

Cambarus diogenes Hay 1895:478 [in part]. Ortmann 1905:123 [in part]. Rhoades 1944:111 [in part]. Eberly 1954:283 [in part]. Brown 1955:62 [in part]. Marlow 1960:229 [in part]. Page 1985:433 [in part]. Page & Mottesi 1995:23 [in part]. Taylor et al. 1996:29 [in part]. Simon 2001:104 [in part]. Taylor et al. 2007:382 [in part]. Taylor & Schuster 2004:80 [in part]. Taylor Schuster & Wylie 2015:66 [in part]. 

Cambarus diogenes diogenes Marlow 1960:233 [in part]. 

Cambarus (Lacunicambarus) diogenes diogenes Hobbs 1969:110 [in part]; 1974:20 [in part]. Bouchard 1972:56 [in part]; 1974:595 [in part]. 

Cambarus (Lacunicambarus) diogenes Hobbs 1989: 24 [in part]. Thoma et al. 2005:334 [in part]. Thoma & Armitage 2008:iii [in part]. 

Cambarus cf. diogenes Glon 2017:55.

 Lacunicambarus aff. diogenes Glon et al. 2018:604 [in part].

Diagnosis. Eyes pigmented, not reduced. Rostrum curved downwards in lateral view, margins converging, slightly thickened, without marginal spines or tubercles, lacking median carina, shallowly excavated. Acumen distinctly delimited basally by 45° angles. Cephalothorax cylindrical, with 3–10 (mean ± sd: 6 ± 1) small tubercles lining posterior margin of cervical groove. Anteroventral branchiostegal tubercles small, numbering 8–29 (mean ± sd: 18 ± 4). Suborbital angle acute. Postorbital ridges developed, lacking anterior spine or tubercle. Areola obliterated, constituting, in adults, 38–45% (mean ± sd: 42 ± 0 %) of entire length of cephalothorax. Antennal scale 2.41 to 3.35 (mean ± sd: 2.80 ± 0.18) times as long as wide, broadest distal to midlength, terminating in small spine, mesial margin forming straight edge. Dorsomesial margin of palm of chelae with 3 rows of tubercles, mesial-most row normally consisting of 6–10 (mean ± sd: 7 ± 1) probolos tubercles, running parallel to second row with 4–9 (mean ± sd: 6 ± 1) probolos tubercles, third row running diagonally from mesial base of palm to lateral dactyl articulation in the form of 5–8 (mean ± sd: 7 ± 1) subprobolos tubercles located in shallow dimples. No tufts of elongated setae at mesial base of fixed finger. Opposable margin of dactyl weakly concave at base. Ratio of dactyl length to palm length 1.78–2.49 (mean ± sd: 2.10 ± 0.16). Dorsomedian longitudinal ridges of dactyl and fixed finger of propodus weakly developed. Dorsolateral impression at base of propodus moderate. Ventral surface of chelae with 0–5 (mean ± sd: 2 ± 1) subpalmar tubercles. Mesial margin of dactyl with 12–33 (mean ± sd: 22 ± 4) prominent tubercles. Ventral surface of carpus with single spine on mesial articular rim, mesial margin with 4–10 (mean ± sd: 7 ± 1) spines of varying sizes. Merus spines numbering 2–9 (mean: 5 ± 2) on ventrolateral margin and 7–16 (mean ± sd: 11 ± 2) on ventromesial margin. Mesial ramus of uropod with distomedian spine not reaching caudal margin. Gonopods of Form I males contiguous at base, with moderately pronounced umbo near midlength of caudal surface; terminal elements consisting of 1) short, tapering, distally truncate central projection lacking subapical notch, shorter than mesial process, directed caudally at approximately 90°, reaching past margin of umbo, 2) mesial process with conical base tipped with protruding finger, directed caudally at approximately 90° and overreaching umbo by noticeable amount and 3) inconspicuous caudal knob sometimes present at caudolateral base of central projection. Hooks on ischium of third pereiopods only. Female with annulus ventralis subquadrangular or kiteshaped, approximately as long as wide, rather deeply embedded in sternum, flexible, with posterior half sclerotized and anterior half mildly pliable.

....

....

Ecological Notes. As mentioned above, Lacunicambarus chimera is a primary burrowing crayfish species. Like other Lacunicambarus species, L. chimera is commonly dug from burrows in fine-grained soils along the floodplains of streams and rivers and in roadside ditches. We have also collected this species in burrows on the banks of manmade ponds and in ditches that were lined with large stones. The chimneys at the mouths of L. chimera burrows are often large and conspicuous, attaining heights of 30 cm or more. These burrows, like those of other primary burrowing crayfishes, provide habitat for many other organisms (e.g., Creaser 1931; Pintor & Soluk 2006; Thoma & Armitage 2008). Glon & Thoma (2017) specifically documented the use of L. chimera burrows as brooding burrows by eastern cicada killer wasps in Pike County, Indiana. 

Little is known about the ecology of L. chimera in situ, but specimens which we have kept in laboratory aquariums have readily consumed a variety of aquarium fish foods, snails, earth worms, and leaf litter from streams, suggesting that this species is an opportunistic omnivore. These specimens were mostly active at night, when they foraged around their enclosures. During the day, they rested inside of artificial burrows made from PVC pipes, occasionally twitching their antennae in response to stimuli. They did not appear to be particularly aggressive, compared to other crayfish species.

Crayfish Associates. We collected the following primary and secondary-burrowing crayfishes from burrows at sites where we found Lacunicambarus chimera: Creaserinus fodiens (Cottle 1863), C. hortoni (Hobbs & Fitzpatrick 1970), Faxonius immunis (Hagen 1870), L. ludovicianus, L. polychromatus, L. aff. polychromatus, Procambarus acutus (Girard 1852), P. clarkii (Girard 1852), P. gracilis (Bundy in Forbes 1876) and P. viaeviridis (Faxon 1914). While sampling for L. chimera, we focused primarily on sampling for burrowing crayfishes and therefore do not have records of the tertiary-burrowing crayfishes that undoubtedly inhabit open water adjacent to L. chimera burrows. 

Etymology. Our choice of the species epithet “chimera” stems from our first encounter with this species. The first specimens that we caught were freshly molted young adults (approximately 30 mm CL). These specimens bore a bright longitudinal gladiate stripe reminiscent of the stripe in L. ludovicianus, L. miltus, and some populations of L. polychromatus. The bright colors on these specimens were similar to those found in L. polychromatus, and the general shape of these specimens was reminiscent of L. diogenes. These features made L. chimera appear to be a chimera of multiple Lacunicambarus species. To honor the nickname given to this species when it was first discovered by Ray Jezerinac and Whitney Stocker, and also as a reference to its impressive size, we suggest the common name “Crawzilla Crawdad.” 

Mael G. Glon, Roger F. Thoma, Marymegan Daly and John V. Freudenstein. 2019.  Lacunicambarus chimera: A New Species of Burrowing Crayfish (Decapoda: Cambaridae) from Illinois, Indiana, Kentucky, and Tennessee. Zootaxa. 4544(4); 451–478.  DOI:  10.11646/zootaxa.4544.4.1

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Crustacea • 2018] Cambarus loughmani • A New Species of Crayfish (Decapoda: Cambaridae) endemic to the Pre-glacial Teays River Valley in West Virginia, USA ---ScRaBBlE

Cambarus loughmani

Foltz, Sadecky, Fetzner & Thoma

in Foltz, Sadecky, Myers, Fetzner, Welsh, Stocker, Glon & Thoma, 2018. 

 DOI: 10.1080/00222933.2018.1557271

Blue Teays Mudbug  ||    facebook.com/WLUCrayfish

ABSTRACT

A new species of crayfish, Cambarus loughmani sp. nov., is described from the preglacial Teays River Valley of Cabell, Kanawha, Lincoln, Mason, and Putnam counties, West Virginia. The species was previously considered to be part of the Cambarus dubius complex. Loughman et al. restricted C. dubius to an orange colour morph found in central and northern portions of the Allegheny Mountains and Appalachian Plateau in central West Virginia, western Maryland, and south-central Pennsylvania. The new species described herein can be distinguished from all other members of Cambarus Erichson, 1846 by a double row of cristiform tubercles on the palm, an open areola with two rows of punctations, and a consistent blue colouration.

KEYWORDS: Appalachian Plateau, Cambarus, crayfish, new species, West Virginia

Figure 3. Dorsal view of a female specimen of Cambarus loughmani collected from the type locality, burrows adjacent to Little Island Creek, Lincoln County, West Virginia, displaying typical life colours for the species.

Photo by Guenter Schuster.

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Cambarus loughmani Foltz, Sadecky, Fetzner and Thoma sp. nov.

 Cambarus dubius Faxon 1884: 114 [in part]. Faxon 1885: 70, pl. 4: fig. 3; pl. 8: figs 7, 7ʹ [in part]; Dewees 1972: 1, figs 1b–l, 2c–h, 3, 4, 5b–h, 6–16 [in part]. 

Cambarus carolinus dubius.–Faxon 1914: 396, 425 [in part]. 

Cambarus (Jugicambarus) dubius.–Hobbs and Bouchard 1973: 62 [in part]; Hobbs 1974: 18, fig. 60 [in part]; Hobbs 1989: 22, fig. 78 [in part]; Jezerinac et al. 1995: 121, fig. 61 [in part]. 

Cambarus aff. dubius. – Loughman et al. 2015: 534 [in part].

Etymology: The authors of this paper name this crayfish in honour of Dr Zachary J. Loughman, assistant professor of biology, West Liberty University. In recent years, prior to the naming of this species, Dr Loughman has served as one of the primary astacological researchers, contributing greatly to our understanding of ecology, taxonomy, conservation and distribution of Appalachian crayfish species. It is fitting that this crayfish be named in his honour, as both he and this crayfish are reclusive, hard to track down, and when faced with adversity never back down and often advance with arms flailing. The authors of this paper are composed of a mix of age classes. For the younger authors, Dr Loughman has served as a teacher, advisor and mentor. For the older authors he has served as a colleague, collaborator and student. For all of us, he has been a friend and an inspiration. Common name:The suggested common name for this species is Blue Teays Mudbug as it inhabits the Teays River Valley.

David A. Foltz II, Nicole M. Sadecky, Greg A. Myers, James W. Fetzner Jr., Stuart A. Welsh, G. Whitney Stocker, Mael G. Glon and Roger F. Thoma. 2018. 

Cambarus loughmani, A New Species of Crayfish (Decapoda: Cambaridae) endemic to the Pre-glacial Teays River Valley in West Virginia, USA. Journal of Natural History.  52(45-46);  DOI: 10.1080/00222933.2018.1557271 

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شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.