We compared Sedum formosanum with related species using morphological traits and molecular phylogenetic analysis of nrITS sequences. Morphological comparisons revealed that the plants historically treated as S. formosanum in the Danjo Islands of Japan had 4-merous flower; 8 stamens; narrow triangular sepals of equal size; horizontal carpels when matured; and an irregular branching form. These traits differed from those of S. formosanum in other regions, which has 5-merous flowers; 10 stamens, thick spatulate sepals of unequal size; erect carpels when matured; and a trichotomous branching form. Phylogenetic analysis indicated that plants known as S. formosanum in the Danjo Islands were sister group to S. tetractinum, which are endemic to China and belong to a different clade than S. formosanum, which are found in other regions. Based on the present morphological comparisons and phylogenetical analyses, we describe plants from the Danjo Islands as a new species, Sedum danjoense, which is distinct from S. formosanum.
Keywords: East Asia, ITS, Phylogeny, Sedum formosanum, Succulent, Eudicots
FIGURE 3. Sedum danjoense Takuro Ito, H. Nakanishi & G. Kokub. A. Habitat and habit. B. Inflorescence. C. Adaxial surface. D. Abaxial surface. E. Flower. F. Sepals. G. Carpels. H. Branching.
Scale bars are 25 mm for A, 5 mm for B–H [A. Wild individuals in Yorishima island photo by Yoshiro Chichibu in May 1989; B. Cultivated in Nagasaki Subtropical Botanical Garden photo by Kiyotaka Minota in Sep. 2011; C-H. Takuro Ito 3658 in Oct. 2016]
Sedum danjoense Takuro Ito, H. Nakanishi & G. Kokub., sp. nov.
Type:— JAPAN. The Kyusyu, the Danjo Islands, Yorishima Island, 25 October 2016 (cultivated in Nagasaki Subtropical Botanical Garden after collecting its natural habit in 1989), Takuro Ito 3658. (holotype TNS!).
Etymology:— The epithet refers to the Japanese name of type locality of the Danjo Islands.
Japanese common name:—Danjo-mannen-gusa (nov.).
Distribution and habitat:— Endemic to the Danjo Islands (Kyusyu), on sunny, coastal rocky slopes exposed to direct sunlight; in typical coastal vegetation within “a community of Miscanthus condensatus Hackel (1899: 639)– Crepidiastrum lanceolatum (Houtt.) Nakai (1920: 150)” similar to those in other regions of Japan, Taiwan and the Philippines. Itow & Nakanishi (1990) mentioned that this species was distributed throughout the Danjo Islands (as S. formosanum), and thus further field surveys are required on the islands.
Takuro Ito, Hiroki Nakanishi, Yoshiro Chichibu, Kiyotaka Minoda and Goro Kokubugata. 2017. Sedum danjoense (Crassulaceae), A New Species of Succulent Plants from the Danjo Islands in Japan. Phytotaxa. 309(1); 23–34. DOI: 10.11646/phytotaxa.309.1.2
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ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
Sedum section Gormania was restricted to Oregon, Nevada and California in the western United States. After extensive field work from 2011 to 2016, we revised 17 members of the group using floral and vegetative characters, resulting in the acceptance of four new taxa in California. A serpentine endemic from the mountains of western Tehama County was recognized asS. rubiginosum. It was separated from S. kiersteadiae by its dense rosettes, overlapping stem leaves and non-apiculate corolla. A serpentine endemic from low elevation canyons in Del Norte County was described as S. patens. It was distinguished from S. laxum by its white spreading petals and yellow anthers. A plant of high elevation, serpentine and non-serpentine sites in Siskiyou County was circumscribed asS. marmorense; it differed from S. oregonense in its sepals and inflorescence with a thick granular waxy deposit, and leaves in dense rosettes. Sedum paradisumwas segregated from S. obtusatum, raised to species level, and divided into two subspecies. Plants of the northern Sierra Nevada were newly defined as S. paradisumsubsp. subroseum, separable with nodding young flowering shoots and a disjunct range in Butte, Plumas and Sierra counties.Sedum flavidumand Sedum eastwoodiae were removed from S. laxum sensu stricto, and raised to species rank, based on floral characters. We clarified the concept of S. obtusatum subsp. retusum, and restored it to the rank of species asS. sanhedrinum; it was restricted to Glenn, Lake, Mendocino, and Tehama counties, California. Sedum flavidum and S. oregonense as defined here showed more morphological variation than previously understood. Finally, we remarked on hybridization and cleistogamy observed in the field.
Keywords: Sedum, new species, California, Eudicots
Figure 10. The petal tips erect and acuminate, essentially parallel to the floral axis. A–D. Sedum laxum subsp. laxum. A–B. Curry Co., Oregon (Wilson & Otting CWG-240). C–D. Josephine Co., Oregon (Zika 25655). E. Sedum moranii, Josephine Co. (Zika 25631). Figure 11. Petal tips narrow in the distal half; the petals strongly spreading to slightly reflexed. A. Sedum citrinum, Del Norte Co., California (Zika 26620 & Brainerd). B. Sedum kiersteadiae, Trinity Co., California (Zika 26294). C. Sedum patens, Del Norte Co. (Zika 26609 & Brainerd). D. Sedum rubiginosum, Tehama Co., California (Zika 25522 & J.K. Nelson). Figure 12. Variation in the width of the distal half of the petal in Sedum flavidum. A. Narrow-petaled extreme, Humboldt Co., California (Wilson 18112). B–D. Variation within a population, Trinity Co. (Zika 25922 & J.K. Nelson). B. Narrow-petaled extreme. C. Slightly narrowed petals. D. Typical broad petals. E. Slightly narrowed petals, Trinity Co., California (CWG-102a). Figure 13. Distal half of the petals broad and ascending. A. Sedum flavidum, Trinity Co., California (J.K. Nelson JKN-12-2). B. Sedum i, Nevada Co., California (Zika 26271). C. Sedum oregonense, Del Norte Co., California (Otting CWG-122). D. Sedum paradisum subsp. i, Plumas Co., California (Wilson, Janeway & Zika CWG-14).
Figure 14. Cultivated individuals often had petals more widely spreading than in the wild. A–B. Sedum flavidum corollas, Humboldt Co., California (Brainerd & Otting CWG-103). A. Ascending petals, wild plants. B. Spreading petals, cultivated plants.C–D. Sedum laxum subsp. laxum corollas, Del Norte Co., California. C. Erect petals, wild plants (Brainerd & Otting CWG-107). D. Ascending petals, cultivated plants (Brainerd & Otting CWG-104). E–F. Sedum sanhedrinum corollas, Tehama Co., California. E. Erect petals, wild plants (Zika 26244). F. Ascending petals, cultivated plants, Tehama Co. (J.K. Nelson JKN-2).
Figure 15. The color of the petals, anthers, filaments, and carpels often changed after anthesis. A. Sedum albomarginatum, after anthesis petals turned orange-brown; Butte Co., California (Zika 25867). B. Sedum flavidum, buds deep pink especially along the keel, faded to white at anthesis, then aged to a uniform deep pink; Humboldt Co., California (Wilson, Otting, & Darington CWG-118). C. Sedum laxum subsp. laxum, after anthesis green carpels turned dark red, red anthers turned white; Del Norte Co., California (Zika 25936). D. Sedum paradisum subsp. subroseum, after anthesis green carpels turned dark red, yellow anthers turned white; Plumas Co., California (Wilson, Zika & Janeway CWG-15). E. Sedum rubiginosum, after anthesis red-based yellow petals turned pallid; Tehama Co., California (Zika 26234).
Figure 16. Some taxa occasionally produced paler flowers in cultivation. A–B. Sedum laxum subsp. laxum. A. Pink petals, wild plants, Del Norte Co., California (Zika 25927). B. Fresh white petals with faint pink background, turning darker pink with age, cultivated, from Josephine Co., Oregon (Zika 25487). C–D. Sedum paradisum subsp. paradisum, Shasta Co., California. C. Yellowish petals, fading pinkish-orange, wild plants (Zika 25924). D. White petals, cultivated (Lindstrand III & Van Susteren NSR-07).
Taxonomic Treatment
1. Sedum albomarginatum Clausen (1975: 424).
2. Sedum citrinum Zika (2014: 112).
3. Sedum eastwoodiae (Britton in Britton & Rose 1903: 31) Berger (1930: 451).
7. Sedum marmorense Otting & R. E. Brainerd, sp. nov.
Species Sedum oblanceolatum proxima sed differt foliis basalibus rosulatis obovatis, a Sedum oregonensis rosulis densissimis et inflorescentiis granularibus ceraceisque.
Etymology:— Sedum marmorense, or Marble Mountains stonecrop, was named for the Marble Mountains and for the marble substrate on which it sometimes grows.
Gormania obtusata (A.Gray) Britton in Britton & Rose (1903: 29).
Cotyledon obtusata (A.Gray) Fedde in Schumann & Fedde (1904: 827).
Echeveriaobtusata (A.Gray) Nelson & Macbride (1913: 476).
Figure 38.Sedum oregonense. A. Habit, with numerous branched sterile shoots, Josephine Co., Oregon (Zika 25657). B–D. Stem leaves, varied from much longer than wide to ± as long as wide, truncated or slightly clasping at base. Colors varied from red to pink, orange, purplish, green or gray. B. Zika 25657. C. Linn Co., Oregon (Zika 25962). D. Clackamas Co., Oregon (Zika 25964). E. Loose rosettes, with easily visible internodes between leaves of the sterile shoots, slightly glaucous, Del Norte Co., California (Otting CWG-122). F. Nodding or arching young inflorescences, pre-anthesis, Lane Co., Oregon (Zika 27948). G–I, K. Inflorescences, varied from flat-topped to capitate or cylindrical. J–K. Flowers, petals typically white, rarely pale yellow, petals ascending, anthers yellow, Josephine Co. G. Del Norte Co. (Zika 25928). H–I, K. Josephine Co. H, K. Zika 25657. I–J. Brainerd & Otting CWG-243.
12a. Sedum paradisum (Denton 1978: 236) Denton ex B. L. Wilson subsp. paradisum.
12b. Sedum paradisum (Denton 1978: 236) Denton ex B. L. Wilson subsp. subroseum B. L. Wilson & Zika, subsp. nov.
Species nostra Sedum paradisum subsp. paradisum aemulans, differt axe inflorescentiae juvenili nutante, a Sedum obtusatum subsp. obtusatum floribus albis postea saepe pallide erubescentibus recedens.
Etymology:— Sedum paradisum subsp. subroseum, or Plumas stonecrop, was named for its flowers, which usually turned pinkish and with age gave the entire inflorescence a rosy appearance.
13. Sedum patens Zika, sp. nov.
Species insignis Sedum laxum eximie affinis sed petalis niveis apicibus patentibus necnon antheris luteis notabilis.
Etymology:— Sedum patens, or Smith River stonecrop, was named for the widely spreading petals.
Figure 43. Sedum rubiginosum, Tehama Co., California. A–C. Habitat, rocky serpentine slopes. A. Photo: J.K. Nelson (Zika 25522 & J.K. Nelson). B. Wilson & Coberly CWG-10. C. Zika 26238. D–E. Flowering shoots (Zika 25522 & J.K. Nelson). F. Dense rosettes (Zika 26234). G. Oblong overlapping ascending stem leaves, faintly glaucous, with truncated bases (Zika 26238). H. Cylindrical inflorescence in early anthesis, flowers dense (Zika 25522 & J.K. Nelson). I–K. Flowers, petals narrow, acute, spreading, reddish towards base, anthers dark red. I. Typical narrow petals (Zika 25522 & J.K. Nelson). J. Unusually broad petals (Zika 26238). K. Relatively short sepals (Zika 26234).
14. Sedum rubiginosum Zika & B. L. Wilson, sp. nov.
Differt haec species a Sedum kiersteadiae rosulis foliorum densis, floribus numerosioribus, sine vel reduci mucrone abaxiali subterminali, necnon foliis caulinis numerosis imbricatisque.
Etymology:— Sedum rubiginosum, or Tedoc stonecrop, was named for the reddish colors found in the fresh and aged petals, as well as the rusty red anthers.
Figure 45. Sedum sanhedrinum. A. Habit, showing large rosette leaves and ascending stem leaves, Mendocino Co., California (Zika 26629 & Brainerd). B–D. Variation in stem leaves, spreading to ascending, broadly to narrowly obovate, bases truncated, slightly glaucous, green to purple or pink. B. Zika 26629 & Brainerd. C–D. Mendocino Co. (Zika 26633 & Brainerd). E. Dense and small rosette leaves, Tehama Co., California (Zika 26244). F. Capitate inflorescence, with dense flowers, Lake Co., California (Wilson & Otting CWG-231); inflorescences were also cylindrical (e.g., Fig. 45D, type) or flat-topped. G–K. Variation in flowers, corolla color pale yellow, orangeyellow, or bright yellow, faded to pink, orange-brown, pale brown, or yellow-brown, white or dull red; anthers yellow or, rarely, dull orange-red, faded to pink, reddish, white, or dull yellow; sepals elongated after anthesis. G. Tehama Co. (Wilson & Otting CWG-234). H. Zika 26629 & Brainerd. I. Zika 26633 & Brainerd. J–K. Petals erect, tips acute to blunt, Tehama Co. (Zika 25970).
15. Sedum sanhedrinum Berger (1930: 451).
Gormania retusa Rose in Britton & Rose (1903: 31).
Cotyledon retusa (Rose) Fedde in Schumann & Fedde (1904: 828).
Sedum laxum (Britton in Britton & Rose 1903: 29) Berger (1930: 451) subsp. retusum (Rose) Clausen (1942: 39).
Peter F. Zika, Barbara L. Wilson, Richard E. Brainerd, Nick Otting, Steve Darington, Brian J. Knaus and Julie Kierstead Nelson. 2018. A Review of Sedum section Gormania (Crassulaceae) in western North America. Phytotaxa. 368(1); 1–61. DOI: 10.11646/phytotaxa.368.1.1
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شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
Pachyphytum (Crassulaceae) is a genus of perennial plants with ca. 20 species endemic to central Mexico. Pachyphytum rogeliocardenasii is described here as a new species. It is compared to P. garciae, which is morphologically similar. The new species is endemic to the northwestern region of the state of Querétaro where it was found on limestone walls. It is assigned the category of endangered (EN).
FIGURE 2. Pachyphytum rogeliocardenasii: A) habit; B) petal color; C) calyx lobes; D) inflorescence peduncle and floral bracts.
FIGURE 2. Pachyphytum rogeliocardenasii: A) habit; B) petal color; C) calyx lobes; D) inflorescence peduncle and floral bracts. P. garciae: E) habit; F) petal color; G) calyx lobes; H) inflorescence peduncle and flowers bracts.
Pachyphytum rogeliocardenasii E. Pérez-Calix & R.Torres, sp. nov.,
Type:— MEXICO. Querétaro: municipio Arroyo Seco, cerca de 2.9 km al sur de Santa María de Cocos, 800 m elevation, 21°18’24.64” N, 99°38’26.54” O, 3 November 2015, E. Pérez 6616 (holotype IEB!, isotypes MEXU!, QMEX!).
Pachyphytum rogeliocardenasii is morphologically similar to P. garciae, which differs in that its leaves are up to 1.3 cm longer and 0.5 cm wider; the bracts of the peduncle are 6‒7.5 mm longer and 2‒3.5 wider; calyx lobes are also larger and corolla lobes are white with an abaxial red spot in the middle.
....
Distribution and habitat:— Pachyphytum rogeliocardenasii is endemic to the northwest region of Queretaro, near the border with Guanajuato where one population that lives on vertical walls of limestone sedimentary rock has been recorded in the canyon of the Atarjea river. The climate of the region is subhumid, semi-warm (sensu García 1973). The vegetation corresponds to the deciduous tropical forest (sensu Zamudio et al. 1992). Some of its elements are Celtis iguanaea and various representatives of Leguminosae. The walls where the plant grows are also habitat by populations of Agave sp., Hechtia sp., Echeveria sp. and Sedum corynephyllum.
Etymology:— The specific epithet is dedicated to the memory of Rogelio Mariano Cárdenas-Soriano (Cuautla, Morelos, 1961 - Pátzcuaro, Michoacán, 2010). Rogelio worked as a scientific illustrator in the Bajío Regional Center of the Institute of Ecology, A.C., from 1995 until his death, and elaborated a series of illustrations, published in the fascicles of Flora del Bajío and of adjacent regions; he also drew new taxa for science, which were published in various specialized journals in botany.
Emmanuel Pérez Calix and Rafael Torres Colín. 2018. Pachyphytum rogeliocardenasii (Crassulaceae), A New Species from northwestern Querétaro, Mexico. Phytotaxa. 348(1); 56–62. DOI: 10.11646/phytotaxa.348.1.7
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شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
