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[Ornithology • 2017] Myzomela irianawidodoae • A Colourful New Species of Myzomela Honeyeater from Rote Island in eastern Indonesia ---ScRaBBlE

Myzomela irianawidodoae Prawiradilaga, Baveja, Suparno, Ashari, Ng, Gwee, Verbelen & Rheindt, 2017  photo:   Philippe Verbelen  e-journ...

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Showing posts with label UK. Show all posts
Showing posts with label UK. Show all posts

Wednesday, March 20, 2019

[Paleontology • 2017] Ieldraan melkshamensis • A New Metriorhynchid Crocodylomorph from the Oxford Clay Formation (Middle Jurassic) of England, with Implications for the Origin and Diversification of Geosaurini ---ScRaBBlE


Ieldraan melkshamensis 
Foffa, Young, Brusatte, Graham & Steel, 2017

Abstract
Metriorhynchids are an extinct group of Jurassic–Cretaceous crocodylomorphs secondarily adapted to a marine lifestyle. A new metriorhynchid crocodylomorph from the Oxford Clay Formation (Callovian, Middle Jurassic) of England is described. The specimen is a large, fragmentary skull and associated single ramus of a lower jaw uniquely preserved in a septarian concretion. The description of the specimen reveals a series of autapomorphies (apicobasal flutings on the middle labial surface of the tooth crowns, greatly enlarged basoccipital tuberosities) and a unique combination of characters that warrant the creation of a new genus and speciesIeldraan melkshamensis gen. et sp. nov. This taxon shares numerous characters with the Late Jurassic–Early Cretaceous genus Geosaurus: tooth crowns that have three apicobasal facets on their labial surface, subtly ornamented skull and lower jaws elements, and reception pits along the lateral margin of the dentary (maxillary overbite). Phylogenetic analysis places this new species as the sister taxon to Geosaurus. The new taxon adds valuable information on the time of origin of the macrophagous subclade Geosaurini, which was initially thought to have evolved and radiated during the Late Jurassic. The presence of Ieldraan melkshamensis, the phylogenetic re-evaluation of Suchodus durobrivensis as a Plesiosuchus sister taxon and recently identified Callovian Dakosaurus-like specimens in the Oxford Clay Formation, indicate that all major Geosaurini lineages originated earlier than previously supposed. This has major implications for the evolution of macropredation in the group. Specifically, we can now demonstrate that the four different forms of true ziphodonty observed in derived geosaurins independently evolved from a single non-functional microziphodont common ancestor.

Keywords: Ieldraan, Melksham monster, Geosaurus, Geosaurini, Jurassic, macrophagy


 Skull and left mandibular ramus of Ieldraan melkshamensis gen. et sp. nov. (NHMUK PV OR 46797).

Systematic palaeontology 
Superorder Crocodylomorpha Hay, 1930 (sensu Walker 1970) 
Suborder Thalattosuchia Fraas, 1901 (sensu Young & Andrade 2009) 
Family Metriorhynchidae Fitzinger, 1843 (sensu Young & Andrade 2009) 
Subfamily Geosaurinae Lydekker, 1889 (sensu Young & Andrade 2009) 
Tribe Geosaurini Lydekker, 1889 (sensu Cau & Fanti 2011)

 Subtribe Geosaurina subtr. nov.

 Type genus. Geosaurus Cuvier, 1824 (sensu Young et al. 2012).

 Geological range. Middle Callovian to Valanginian (34 myr duration). 
Geographical range. European endemic (UK, Germany and France). 

 Genus Ieldraan gen. nov. 
Type species. Ieldraan melkshamensis gen. et sp. nov. 

 Derivation of the name: Older One’. Ieldra, Old English for older; and an, Old English for one, referring to the stratigraphically older age of this new genus compared to its close relative Geosaurus

Ieldraan melkshamensis sp. nov.  
1888 Metriorhynchus moreli Eudes-Deslongchamps; Lydekker: 97.

Derivation of name: ‘Older One from Melksham’, epithet translated from Latin, locative case.


Conclusions:
Based on our description of a long overlooked and misinterpreted specimen (NHMUK PV OR 46797), we establish the new taxon Ieldraan melkshamensis gen. et sp. nov. Despite the poor state of preservation, we demonstrate that this late Middle Jurassic taxon from the OCF shows remarkable similarities with the Late Jurassic genus Geosaurus. Ieldraan and Geosaurus are found to be sister taxa in a new European endemic, Callovian–Valanginian geosaurin lineage that we name Geosaurina subtr. nov. The morphology and stratigraphical occurrence of Ieldraan melkshamensis, combined with our phylogenetic analysis, demonstrate that numerous adaptations linked to macrophagy had already evolved in Geosaurini by the Callovian stage. This suggests that the diversification of the tribe was perhaps less abrupt than previously thought, but rather had a longer temporal and phylogenetic fuse. We also show that the evolution of ziphodonty followed a different path than previously hypothesized. The new information presented here indicates that four different true ziphodont morphologies in the derived Late Jurassic geosaurins independently evolved from a unique non-functional microziphodont common ancestor.


 Davide Foffa, Mark T. Young, Stephen L. Brusatte, Mark R. Graham and Lorna Steel. 2017. A New Metriorhynchid Crocodylomorph from the Oxford Clay Formation (Middle Jurassic) of England, with Implications for the Origin and Diversification of Geosaurini. Journal of Systematic Palaeontology.  DOI: 10.1080/14772019.2017.1367730

  

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[PaleoMammalogy • 2017] Durlstodon ensomi & Durlstotherium newmani • Highly Derived Eutherian Mammals from the earliest Cretaceous of southern Britain ---ScRaBBlE


the Purbeck lagoon at dusk with Durlstodon gen. nov. (left foreground), Durlstotherium gen. nov. (right and center foreground) and the theropod Nuthetes holding a captured Durlstotherium (centre middle distance). 

A. Durlstotherium newmani gen. et sp. nov., NHMUK PV M 99991.
B. Durlstodon ensomi gen. et sp. nov.
, NHMUK PV M 99992.

Artwork by Mark Witton. @MarkWitton 

Eutherian mammals (Placentalia and all mammals phylogenetically closer to placentals than to marsupials) comprise the vast majority of extant Mammalia. Among these there is a phenomenal range of forms and sizes, but the origins of crown group placentals are obscure. They lie within the generally tiny mammals of the Mesozoic, represented for the most part by isolated teeth and jaws, and there is strongly conflicting evidence from phenomic and molecular data as to the date of origin of both Eutheria and Placentalia. The oldest purported eutherians are Juramaia from the Upper Jurassic of China, and Eomaia and Acristatherium from the Lower Cretaceous, also of China. Based on dental characters and analyses of other morphological and molecular data, doubt has recently been cast on the eutherian affinities of the Chinese taxa and consequently on the date of emergence of Eutheria. Until now, the only tribosphenic mammal recorded from the earliest Cretaceous (Berriasian) Purbeck Group of Britain was the stem tribosphenidan Tribactonodon. Here we document two new tribosphenic mammals from the Purbeck Group, Durlstotherium gen. nov. and Durlstodon gen. nov., showing highly derived eutherian molar characters that support the early emergence of this clade, prior to the Cretaceous.

Key words: Mammalia, Eutheria, dentition, Early Cretaceous, Purbeck Group, Britain.


Fig. 5. Normal light photographs of studied specimens of eutherian mammal teeth from the Berriasian Purbeck Group of Dorset, southern England; in mesial (A1, B1), distal (A2, B2), lingual (A3, B3), and labial (A4, B4) views.
 A. Durlstotherium newmani gen. et sp. nov., NHMUK PV M 99991. B. Durlstodon ensomi gen. et sp. nov., NHMUK PV M 99992.

Systematic palaeontology
Class Mammalia Linnaeus, 1758
Subclass Tribosphenida McKenna, 1975
Infraclass Eutheria Gill, 1872
Order and Family incertae sedis

Genus Durlstotherium nov.

Type species: Durlstotherium newmani sp. nov., monotypic, see below.

Etymology: In reference to Durlston Bay, Dorset, UK, the locality from where the holotype and only specimen was obtained; and from Greek, therion, beast; a commonly used suffix in the names of mammals.; In honour of Charlie Newman, recognising his intimate local knowledge, willingly shared, and his help in the field.


Genus Durlstodon nov. 
Type species: Durlstodon ensomi sp. nov., monotypic, see below.

Etymology: In reference to Durlston Bay, Dorset, UK, the locality from where the holotype and only specimen was obtained; and from Greek, odontos, tooth.; In honour of Paul Ensom, recognising his major contribution to Purbeck palaeontology.

Fig. 7. Artist’s impression of the Purbeck lagoon at dusk with Durlstodon gen. nov. (left foreground), Durlstotherium gen. nov. (right and center foreground) and the theropod Nuthetes holding a captured Durlstotherium (centre middle distance).

Artwork by Mark Witton. @MarkWitton

Steven C. Sweetman, Grant Smith and David M. Martill. 2017. Highly Derived Eutherian Mammals from the earliest Cretaceous of southern Britain. Acta Palaeontologica Polonica. in press. DOI: 10.4202/app.00408.2017


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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Thaumatodracon wiedenrothi • A Morphometrically and Stratigraphically Intermediate New Rhomaleosaurid Plesiosaurian from the Lower Jurassic (Sinemurian) of Lyme Regis, England ---ScRaBBlE


Thaumatodracon wiedenrothi  Smith & Araújo, 2017

Illustration: L. Soares. DOI: 10.1127/pala/308/2017/89  

Abstract

An excellently preserved partial skeleton of a rhomaleosaurid plesiosaurian (NLMH 106. 058) from the Sinemurian (Lower Jurassic) of Lyme Regis, England, is described. The material consists of a complete cranium, mandible, and articulated cervical vertebral column. It is noteworthy because large-headed rhomaleosaurids are rare from this stratigraphic horizon and it is taxonomically distinct. The material is referred to a new taxonThaumatodracon wiedenrothi gen. nov. et sp. nov, diagnosed by two autapomorphies: 1. a pronounced transverse trough on the posterior margin of the dorsal ramus of the squamosal; 2. possibly paired anteriorly tapering triangular basioccipital processes. It also possesses a unique combination of other characters including a ‘short’ premaxillary rostrum (length and width subequal), five premaxillary alveoli, premaxilla-maxilla sutures parallel anterior to the external nares, frontals contact on the midline, prefrontal-frontal suture convex and gently curved medially, mandibular symphyseal region spatulate and ‘short’ (length and width subequal), prominent dorsally concave medial flange anteromedial to the articular glenoid, robust rod-like axis neural spine with a circular transverse cross section, and cervical neural spines with a mediolaterally expanded apex. The taxon shares some of these characters with earlier Hettangian rhomaleosaurids (e. g. Atychodracon, Eurycleidus), and other characters with later Toarcian rhomaleosaurids (e. g. Rhomaleosaurus sensu stricto and Meyerasaurus). Inclusion of Thaumatodracon as an additional operational taxonomic unit in several existing cladistic analyses demonstrates that it occupies a relatively derived position within Rhomaleosauridae. A morphometric multivariate analysis of Lower Jurassic rhomaleosaurids shows that Thaumatodracon is also proportionally intermediate between known rhomaleosaurid taxa. Thaumatodracon is therefore a stratigraphically and anatomically intermediate taxon that fills a gap in our knowledge of the evolution of this macro-predatory plesiosaurian clade.

Keywords: Plesiosauria, Sauropterygia, Rhomaleosauridae, Lower Jurassic, Lyme Regis


Thaumatodracon wiedenrothi
Illustration: Luzia Soares. 

 Adam S. Smith and Ricardo Araújo. 2017. Thaumatodracon wiedenrothi, A Morphometrically and Stratigraphically Intermediate New Rhomaleosaurid Plesiosaurian from the Lower Jurassic (Sinemurian) of Lyme Regis. Palaeontographica, Abt. A: Palaeozoology – Stratigraphy 4-6; 89 - 125.  DOI: 10.1127/pala/308/2017/89  

 Adam S. Smith and Ricardo Araújo. 2017. Morphometric data and phylogenetic analysis of Thaumatodracon wiedenrothiPANGAEA. DOI: 10.1594/PANGAEA.870543

   


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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] On the Largest Ichthyosaurus: A New Specimen of Ichthyosaurus somersetensis Containing An Embryo ---ScRaBBlE


Ichthyosaurus somersetensis Lomax and Massare, 2017
 (NLMH 106234) 
from the Lower Jurassic (lower Hettangian) of Doniford Bay, Watchet, Somerset, UK


A formerly undescribed Ichthyosaurus specimen from the collection of the Niedersächsisches Landesmuseum (Lower Saxony State Museum) in Hannover, Germany, provides valuable new information. The skeleton was collected from the Lower Jurassic strata (lower Hettangian, Blue Lias Formation) of Doniford Bay, Somerset, UK. However, the specimen is a composite as almost the entire tail has been added and other parts are reconstructed. Regardless of the incomplete preservation, the estimated total length of this individual, based on the skull and precaudal length, is between 300 and 330 cm and it is thus the largest unequivocal example of the genus Ichthyosaurus. Cranial and postcranial characters, specifically from the maxilla, lacrimal, jugal, the humerus, and the ilium justify a referral to I. somersetensis. A fork-like shape of the proximal end of the ilium is unusual and has not been reported for any species of Ichthyosaurus. Likewise the presence of four elements in the third row of the hindfin, indicated by the presence of a bifurcation is novel for the species and has wider implications for the taxonomic utility of hindfins within the genus. The specimen also bears an embryo, which is only the third embryo known for Ichthyosaurus and the first to be positively identified to species level.

Key words: Ichthyosauria, Ichthyosaurus somersetensis, embryo, Jurassic, Hettangian, UK, Somerset.


Artist impression of pregnant Ichthyosaurus.
ILLUSTRATION: Joschua Knüppe

Fig. 2.  Skeleton of Ichthyosaurus somersetensis Lomax and Massare, 2017 (NLMH 106234) from the Lower Jurassic (lower Hettangian) of Doniford Bay, Watchet, Somerset, UK. 


Systematic palaeontology

Order Ichthyosauria de Blainville, 1835
Family Ichthyosauridae Bonaparte, 1841
Genus Ichthyosaurus De la Beche and Conybeare, 1821
Type species: Ichthyosaurus communis De la Beche and Conybeare, 1821; upper Hettangian–lower Sinemurian, Lower Jurassic of England, UK.

Ichthyosaurus somersetensis Lomax and Massare, 2017

 .....


Dean R. Lomax and Sven Sachs. 2017.  On the Largest Ichthyosaurus: A New Specimen of Ichthyosaurus somersetensis Containing An Embryo. Acta Palaeontologica Polonica. in press.  DOI: 10.4202/app.00376.2017

Largest 'Sea Dragon' Fossil Accidentally Discovered in Museum
on.natgeo.com/2wNrj9z   @NatGeo


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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Cascolus ravitis • A New Crustacean from the Herefordshire (Silurian) Lagerstätte, UK , and Its Significance in Malacostracan Evolution ---ScRaBBlE


Cascolus ravitis 
Siveter, Briggs, Siveter, Sutton & Legg, 2017  


Abstract

Cascolus ravitis gen. et sp. nov. is a three-dimensionally preserved fossil crustacean with soft parts from the Herefordshire (Silurian) Lagerstätte, UK. It is characterized by a head with a head shield and five limb pairs, and a thorax (pereon) with nine appendage-bearing segments followed by an apodous abdomen (pleon). All the appendages except the first are biramous and have a gnathobase. The post-mandibular appendages are similar one to another, and bear petal-shaped epipods that probably functioned as a part of the respiratory–circulatory system. Cladistic analysis resolves the new taxon as a stem-group leptostracan (Malacostraca). This well-preserved arthropod provides novel insights into the evolution of appendage morphology, tagmosis and the possible respiratory–circulatory physiology of a basal malacostracan.

An international team of scientists led by the University of Leicester has discovered a new 430 million-year-old fossil and has named it in honour of Sir David Attenborough - who grew up on the University campus.  

Systematic palaeontology

Phylum Arthropoda
Subphylum Crustacea

Class Malacostraca
Subclass Phyllocarida

Order Leptostraca

Genus Cascolus gen. nov.

Type species: Cascolus ravitis sp. nov.

Etymology: The new crustacean is named in honour of the naturalist and broadcaster Sir David Attenborough, who grew up on University College Leicester campus, in celebration of his 90th birthday. Latin castrum stronghold’ and colus dwelling in’; alluding to the Middle/Old English source for the surname ‘Attenborough, derived from attenat the’ and burgh a fortified place’. Latin Ratae, the Roman name for Leicester, vita life’ and commeatis ‘a messenger’.


  Diagnosis of genus (monotypic) and species: An elongate body comprising a head with a head shield, pedunculate eyes and five limb pairs; and a trunk consisting of a thorax (pereon) with nine limb-bearing segments and an apodous abdomen (pleon). The first appendage is uniramous and has three slender flagella longer than the body. All other appendages are biramous and have a gnathobase. The post-mandibular appendages are similar to one another, except that the fourth head appendage bears a single petal-shaped epipod, and the fifth head appendage and each trunk appendage bear two petal-shaped epipods.

  Material: Only known from the holotype OUMNH C.29698 (figure 1v), a specimen with soft parts reconstructed in three dimensions (figure 1a–u).

  Locality and horizon: Herefordshire, England, UK; Wenlock Series, Silurian.


David J. Siveter, Derek E. G. Briggs, Derek J. Siveter, Mark D. Sutton and David Legg. 2017. A New Crustacean from the Herefordshire (Silurian) Lagerstätte, UK, and Its Significance in Malacostracan Evolution. Proceedings of the Royal Society B.  DOI: 10.1098/rspb.2017.0279 

 430 million-year-old fossil named in honor of Sir David Attenborough
 http://phy.so/409322871   @physorg_com

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Lemmysuchus obtusidens • Re-description of ''‘Steneosaurus’ obtusidens'' Andrews, 1909, An Unusual Macrophagous Teleosaurid Crocodylomorph from the Middle Jurassic of England ---ScRaBBlE


Lemmysuchus Steneosaurus obtusidens'' Andrews, 1909


Abstract
Teleosaurids were a clade of crocodylomorphs that attained near-global distribution during the Jurassic Period. Within Teleosauridae, one particular sub-clade of durophagous/macrophagous taxa achieved large body sizes and were apex predators in shallow marine environments during the Late Jurassic and Early Cretaceous in Europe and around the coast of the Tethys Seaway. Unfortunately, the origins of this clade are still poorly understood. ‘Steneosaurus’ obtusidens is a little-studied macrophagous species from the Oxford Clay Formation (Callovian, Middle Jurassic) of the UK and near Migné-les-Lourdines (Middle Callovian) in France. Despite being considered a sister taxon of the Late Jurassic taxon Machimosaurus, the taxonomy of ‘S.’ obtusidens remains unclear. Although three different synonymies have been proposed (variously a subjective synonym of other taxa), these taxonomic hypotheses have not been based on detailed anatomical comparisons and thus have not been tested. Here, we re-describe the holotype of ‘S.’ obtusidens, demonstrate that it is indeed a valid taxon, restrict the referred specimens to a fragmentary skeleton, nearly complete skull, and partial rostrum, and establish a new monotypic genus, Lemmysuchus. Our re-description reveals five autapomorphies for Lemmysuchus obtusidens and nine apomorphic characters that support the tribe Machimosaurini (Lemmysuchus Machimosaurus).

Keywords: Crocodylomorpha, LemmysuchusMachimosaurusSteneosaurus, Teleosauridae, Thalattosuchia




Michela M. Johnson, Mark T. Young, Lorna Steel, Davide Foffa, Adam S. Smith, Stéphane Hua, Philipe Havlik, Eliza A. Howlett and Gareth Dyke. 2017. Re-description of ''‘Steneosaurus’ obtusidens'' Andrews, 1909, An Unusual Macrophagous Teleosaurid Crocodylomorph from the Middle Jurassic of England. Zoological Journal of the Linnean Society. in press.  DOI: 10.1093/zoolinnean/zlx035 



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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2019] Descriptive Anatomy of the Largest Known Specimen of Protoichthyosaurus prostaxalis (Reptilia: Ichthyosauria) including Computed Tomography and Digital Reconstruction of A Three-dimensional Skull ---ScRaBBlE


Protoichthyosaurus prostaxalis Appleby, 1979

in Lomax, Porro & Larkin, 2019.  

Abstract
Ichthyosaur fossils are abundant in Lower Jurassic sediments with nine genera found in the UK. In this paper, we describe the partial skeleton of a large ichthyosaur from the Lower Jurassic (lower Sinemurian) of Warwickshire, England, which was conserved and rearticulated to form the centrepiece of a new permanent gallery at the Thinktank, Birmingham Science Museum in 2015. The unusual three-dimensional preservation of the specimen permitted computed tomography (CT) scanning of individual braincase elements as well as the entire reassembled skull. This represents one of the first times that medical imaging and three-dimensional reconstruction methods have been applied to a large skull of a marine reptile. Data from these scans provide new anatomical information, such as the presence of branching vascular canals within the premaxilla and dentary, and an undescribed dorsal (quadrate) wing of the pterygoid hidden within matrix. Scanning also revealed areas of the skull that had been modelled in wood, clay and other materials after the specimen’s initial discovery, highlighting the utility of applying advanced imaging techniques to historical specimens. Additionally, the CT data served as the basis for a new three-dimensional reconstruction of the skull, in which minor damage was repaired and the preserved bones digitally rearticulated. Thus, for the first time a digital reconstruction of the skull and mandible of a large marine reptile skull is available. Museum records show the specimen was originally identified as an example of Ichthyosaurus communis but we identify this specimen as Protoichthyosaurus prostaxalis. The specimen features a skull nearly twice as long as any previously described specimen of P. prostaxalis, representing an individual with an estimated total body length between 3.2 and 4 m.




Figure 1: Three-dimensional skull of BMT 1955.G35.1, Protoichthyosaurus prostaxalis.
 (A) Original photograph of the first skull reconstruction (left lateral view) within a couple of years of the 1955 excavation. Note that the prefrontal and postorbital are present, which we have been unable to locate in our study. (B) Skull in left lateral view, as reconstructed in 2015. (C) Skull in right lateral view, as reconstructed in 2015. Note the distinctive asymmetric maxilla with long, narrow anterior process. Teeth are not in their original positions. Scale bar represents 20 cm.



Conclusions: 
In this study, we describe a large, partial ichthyosaur skeleton from the Early Jurassic of Warwickshire, England. In addition to examining the specimen, we carried out CT scanning of individual skull bones as well as the entire, reassembled skull, one of the first times the skull of a large marine reptile has been successfully CT-scanned, visualized and reconstructed in 3D (see McGowan, 1989; Foffa et al., 2014a). CT scanning contributed greatly to our anatomical description by revealing features not visible on original fossil material such as: branching, longitudinal vascular canals within the premaxilla and dentary; short canals penetrating the nasal, lacrimal, stapes and articular; trabecular bone within the opisthotic; canals in the basisphenoid and supraoccipital; the presence of the quadrate process of the pterygoid; and the sutural morphology. We also demonstrate the utility of applying medical imaging techniques to historic specimens to differentiate between original fossil material and reconstructed regions, as well as the advantage of using digital visualization to accurately reconstruct large fossil specimens in 3D.

The detailed description of the three-dimensional skull and braincase presented herein also provides information that can be used in phylogenetic studies. Although incomplete, the skull and braincase preserve various elements that have not previously been reported or described in any specimen of Protoichthyosaurus and therefore it provides more information about this taxon so that its phylogenetic position can be explored in more detail. Furthermore, our study has found additional characters that may lend further support for the distinction of Protoichthyosaurus from its sister taxon Ichthyosaurus, such as the morphology of the pterygoid and anteroventral surface of the parietal, which differ from that described for Ichthyosaurus (McGowan, 1973). However, considering that only a couple of specimens preserve these elements, it is possible that the differences may be the result of individual variation; more three-dimensional specimens of both taxa are needed to test and clarify these findings.

Based on a unique combination of characters, we identify the studied specimen as P. prostaxalis. With a skull nearly twice as long as any previously described specimen of P. prostaxalis, this specimen greatly increases the known size range of this genus. Compared with known, contemporaneous Sinemurian ichthyosaurs, the estimated size suggests it was larger than all species of Ichthyosaurus (Lomax & Sachs, 2017), and comparable with the largest known specimens of Leptonectes tenuirostris (McGowan, 1996a), but smaller than L. solei (McGowan, 1993), Excalibosaurus costini (McGowan, 2003) and Temnodontosaurus platyodon (McGowan, 1996b). Thus, our study also provides new information on ichthyosaur diversity and potential ecology in the Early Jurassic of the UK.


Dean R. Lomax, Laura B. Porro and Nigel R. Larkin. 2019. Descriptive Anatomy of the Largest Known Specimen of Protoichthyosaurus prostaxalis (Reptilia: Ichthyosauria) including Computed Tomography and Digital Reconstruction of A Three-dimensional Skull.   PeerJ. 7:e6112. DOI:  10.7717/peerj.6112


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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

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