ScRaBBlE: Pycnonotidae

Showing posts with label Pycnonotidae. Show all posts

Wednesday, March 20, 2019

[Ornithology • 2018] Nok, gen. nov. • Phylogenetic Affinities of the Enigmatic Bare-faced Bulbul Pycnonotus hualon with Description of A New Genus ---ScRaBBlE

Nok hualon (Woxvold, Duckworth & Timmins, 2009)

in Fuchs, Pasquet, Stuart, Woxvold, Duckworth & Bowie, 2018.

DOI: 10.1111/ibi.12580

Abstract

The recently described Bare-faced Bulbul Pycnonotus hualon from Lao PDR has a very distinct morphology and habitat (karsts). Mitochondrial and nuclear data from the type material demonstrated that P. hualon is sister to members of the genus Spizixos. To highlight its unique morphology and phylogenetic distinctiveness, we describe a new monotypic genus for the Bare-faced Bulbul.

The Bare-faced Bulbul Pycnonotus hualon Woxvold, Duckworth & Timmins, 2009, was first collected from the small limestone karst outcrop of Pha Lom, north-east Savannakhet Province, Lao PDR, in late 2008, following a 1995 sighting of strange bald-looking bulbuls about 185 km away, above the Hinboun plain at the northern end of the Khammouan limestone massif. As of 2017, the species remains known only from the karst landforms of Lao PDR between the Pha Lom and above the Hinboun plain. This discovery forms part of a sustained pulse of discoveries of new bird and mammal species in and around the Annamite Mountains of Lao PDR and Vietnam, mostly associated with two distinct habitat types, karst limestone and wet evergreen forest. Several of the newly discovered species are phylogenetically highly distinct. These include a forest bovid (Saola Pseudoryx nghetinhensis Vu, Pham, Nguyen, Do, Arctander & MacKinnon, 1993; Vu et al. 1993); the first representative of Caudata for Lao PDR (Lao Warty Newt Laotriton laoensis (Stuart & Papenfuss, 2002); Stuart & Papenfuss 2002); and a living member of the Diatomyidae (Kha-nyou Laonastes aenigmamus Jenkins, Kilpatrick, Robinson & Timmins, 2005; Jenkins et al. 2005), a lineage of rodent formerly presumed to have been extinct for the past 11 million years (Dawson et al. 2006). Here we investigated the phylogenetic relationships of the Bare-faced Bulbul (Fig. 1).

Figure 1. Image of Bare-faced Bulbul “Pycnonotus hualon”. captured on the Pha Lom limestone massif, northeast Savannakhet Province, Lao PDR in December 2008.

(Photo: I. Woxvold).

by Risso Romain commons.wikimedia.org

Nok, gen. nov.

Type species: Pycnonotus hualon Woxvold, Duckworth and Timmins, 2009 by original description.

Included species: Nok hualon (Woxvold, Duckworth & Timmins, 2009), comb. nov.

Etymology: Nok means bird in Lao (ນົກ) and is used to highlight the range-restricted distribution of the species. Nouns in the Lao language lack gender. Nok is proposed here as a masculine noun.

ນົກຂວກຫົວລ້ານ / ນົກຂວາກຫົວລ້ານ

Jérôme Fuchs, Eric Pasquet, Bryan L. Stuart, Iain A. Woxvold, J. W. Duckworth and Rauri C.K. Bowie. 2018. Phylogenetic Affinities of the Enigmatic Bare-faced Bulbul Pycnonotus hualon with Description of A New Genus. Ibis. The International Journal of Avian Science. DOI: 10.1111/ibi.12580

ResearchGate.net/publication/322697999_Phylogenetic_affinities_Pycnonotus_hualon

facebook.com/bcst.or.th/photos/1593960444052226

I.A. Woxvold; J.W. Duckworth and R.J. Timmins. 2009. An Unusual New Bulbul (Passeriformes: Pycnonotidae) from the Limestone Karst of Lao PDR. Forktail. 25: 1–12. http://lad.nafri.org.la/fulltext/2136-0.pdf

wwf.se/press/pressrum/pressmeddelanden/1307077-nya-sensationella-arter-upptackta-i-sydostasien

ຈາວລໍ່ ຢົງເຢ້. 2014. ການກະຈາຍ ແລະພື້ນທີ່ອາໄສຂອງນົກຂວາກຫົວລ້ານໃນເຂດພູຫີນປູນ ກໍລະນີສຶກສາ: ບ້ານມ່ວງດ່ອຍ, ເມືອງທ່າແຂກ, ແຂວງຄຳມ່ວນ (Distribution and Habitat of Pycnonotus hualon in Limestone Karst: A Case Study in Mouagdoy Village Thakhed District, Khammouane Province). ມະຫາວິທະຍາໄລແຫ່ງຊາດ.
http://dspace.nuol.edu.la:8080/jspui/handle/123456789/1091

---------------------------------------------------------------

روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Ornithology • 2017] Phylogeography of Bulbuls in the Genus Iole (Aves: Pycnonotidae) ---ScRaBBlE

S. Manawatthana, P. Laosinchai, N. Onparn, W.Y. Brockelman & P.D. Round. 2017.
Phylogeography of Bulbuls in the Genus Iole (Aves: Pycnonotidae).

Biol. J. Linn. Soc. 120(4); 931-944. DOI: 10.1093/biolinnean/blw013

Abstract

Southeast Asia is one of the most geologically dynamic regions of the world with great species diversity and high endemism. We studied the bulbuls of the south and southeast Asian genus Iole (Aves: Pycnonotidae) in order to analyse their evolutionary relationships and describe their patterns of diversification and delimit species boundaries. Our phylogeographic reconstruction, based on two mitochondrial and one nuclear markers, sampled from all 13 recognized Iole taxa, presently grouped as four species, revealing three primary lineages: (1) a Palawan lineage (2) a Sundaic group distributed in the Malay Peninsula, Sumatra and Borneo and (3) an Indochinese group distributed throughout continental Southeast Asia. Divergence time estimation suggested that the Palawan lineage diverged during the Miocene (around 9.7 Mya), a later split between the Sundaic and Indochinese lineages occurring around 7.2 Mya. The present classification of Iole based on morphology does not accurately reflect taxonomic relationships within the genus, in which we recognize five more putative species. An integrative approach that incorporates morphology and bioacoustics should further refine our understanding of species limits among Iole taxa.

Keywords: Iole, phylogeny, phylogeography, pycnonotidae, species tree

Sontaya Manawatthana, Parames Laosinchai, Nuttaphon Onparn,Warren Y. Brockelman and Philip D. Round. 2017. Phylogeography of Bulbuls in the Genus Iole (Aves: Pycnonotidae). Biol. J. Linn. Soc. 120(4); 931-944.. DOI: 10.1093/biolinnean/blw013

facebook.com/photo.php?fbid=1471650692853617

สรุปคร่าวๆเฉพาะส่วนของประเทศไทย

นกปรอดสกุล Iole (อ่านว่า ไอโอลี่) เดิมพบได้ในไทย 3 ชนิดคือ

1.) นกปรอดเล็กตาขาว (Grey-eyed Bulbul; Iole propinqua)

2.) นกปรอดเล็กสีไพลตาแดง (Olive Bulbul; Iole viridescens)

3.) นกปรอดหงอนตาขาว (Buff-vented Bulbul; Iole charlottae)

ทั้งสามชนิดมีลักษณะภายนอกใกล้เคียงกันมาก

ซึ่งผลจากการศึกษานี้ทำให้มีนกชนิดใหม่ของไทยเพิ่มขึ้นมาอีก 1 ชนิด คือ

4.) Baker’s Bulbul (Iole cinnamomeoventris; ยังไม่มีชื่อภาษาไทยอย่างเป็นทางการ)

- นกชนิดใหม่นี้เดิมถูกจัดเป็นเพียงชนิดย่อยของนกปรอดเล็กตาขาว เนื่องจากมีลักษณะภายนอกใกล้เคียงปรอดเล็กตาขาวมากๆ จนหลายๆครั้งในภาคสนามไม่เห็นความแตกต่าง

- แต่จากการวิเคราะห์ DNA พบว่ามันมีความแตกต่างในเชิงวิวัฒนาการมากจนสมควรจะแยกเป็นชนิดใหม่ และยังพบว่ามีสายวิวัฒนาการใกล้ชิดกับ ปรอดเล็กสีไพลตาแดง (แทนที่จะใกล้ชิดกับปรอดเล็กตาขาวอย่างที่ถูกจัดไว้เป็นชนิดย่อยเดิม)

- พบในภาคใต้ของไทย บริเวณด้ามขวานตั้งแต่ราวๆ จ.เพชรบุรีลงไปถึงราวๆ จ.สงขลา

- ลักษณะที่ใช้จำแนกในภาคสนามได้ดีที่สุดคือเสียงร้อง จะต่างกับปรอดหงอนตาขาวที่ลักษณะภายนอกคล้ายคลึงกันและพบในบริเวณภาคใต้เหมือนกัน

ตัวอย่างเสียงของ Iole cinnamomeoventris http://www.xeno-canto.org/336909

ตัวอย่างเสียงของ ปรอดหงอนตาขาว Iole charlottae http://www.xeno-canto.org/336936

- ชื่อ Baker's Bulbul ตั้งเพื่อเป็นเกียรติแก่ E. C. Stuart Baker นักปักษีวิทยาชาวอังกฤษ ซึ่งเป็นคนแรกที่จำแนกชนิดย่อย cinnamomeoventris นี้ไว้เมื่อปี 1917

--------------------------------

ถ้านับรวมทั้งเอเชียตะวันออกเฉียงใต้ เดิมนกปรอดสกุลนี้มีทั้งหมด 4 ชนิด ผล DNA จากการศึกษานี้บ่งชี้ว่าควรจะจำแนกเพิ่มอีก 5 ชนิด รวมเป็น 9 ชนิด ใครสนใจสามารถอ่านรายละเอียดได้ในงานวิจัยฉบับเต็มครับ: academic.oup.com/biolinnean/article-abstract/120/4/931/2970060/Phylogeography-of-bulbuls-in-the-genus-Iole-Aves

ทั้งนี้ เปเปอร์แรกนี้จะเน้นเฉพาะผลจากการวิเคราะห์ DNA อาจจะอ่านยากสำหรับคนทั่วไปซักหน่อย ยังมีเปเปอร์ที่สองที่จะพูดถึงลักษณะภายนอก สีขน และผลการวิเคราะห์เสียงร้อง ที่ตอนนี้กำลังอยู่ระหว่างเตรียมส่งตีพิมพ์ครับ

facebook.com/photo.php?fbid=1471650692853617

---------------------------------------------------------------

روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

[Ornithology / Conservation • 2017] Significance of the Globally Threatened Straw-headed Bulbul (Pycnonotus zeylanicus) Populations in Singapore: A Last Straw for the Species? ---ScRaBBlE

Straw-headed Bulbul Pycnonotus zeylanicus

photo: Lee Tiah Khee

SingaporeBirdGroup.wordpress.com BirdLife.org

blog.journals.cambridge.org DOI: 10.1017/S0959270917000028

Abstract

The globally threatened Straw-headed Bulbul Pycnonotus zeylanicus is one of South-East Asia’s most imperiled songbirds due to the surging demand for the species in the regional bird trade. Recently uplisted from Vulnerable to Endangered, populations of the Straw-headed Bulbul have been extirpated from Java, Thailand and possibly Sumatra while those in Borneo and Peninsular Malaysia are in decline. Intriguingly, a significant yet rarely documented population of this species persists in Singapore. A major stronghold in Singapore is Ubin Island where a population is known since the 1920s. Using a long-term citizen science dataset rarely available for South-East Asian bird species, we determined the status and population trends of the Straw-headed Bulbul in Singapore over a 10–15 year period using Poisson regression models and standardised population indices. We found that the Straw-headed Bulbul population has increased at a rate of 3.69 ± 1.21% per annum on Ubin Island, while the population on Singapore Island remained stable (0.56% per annum) from 2000 to 2016. The population trends in Singapore contrast starkly with the declines reported elsewhere in South-East Asia. We estimated the population in Singapore to be a minimum of 202 individuals, distributed over multiple forest patches. The largest subpopulation of about 110 adult individuals persists on Ubin and which alone forms between 6.5–18.3% of the estimated global population in 2016. Given this unique situation, we recommend a number of conservation measures for the Straw-headed Bulbul to better protect the species, including: (1) an expansion of the protected area network in Singapore to include Ubin as a reserve, (2) the development of an endangered species management plan and, (3) the establishment of ex-situ conservation programmes in zoological institutions and wildlife centres in the region.

Straw-headed Bulbul Pycnonotus zeylanicus

photo: Francis Yap

Ding Li Yong, Kim Seng Lim, Kim Chuah Lim, Trixie Tan, Siyang Teo and Hua Chew Ho. 2017. Significance of the Globally Threatened Straw-headed Bulbul (Pycnonotus zeylanicus) Populations in Singapore: A Last Straw for the Species?
Bird Conservation International. DOI: 10.1017/S0959270917000028

ResearchGate.net/publication/309423215_Significance_of_the_globally_threatened_Straw-headed_Bulbul_Pycnonotus_zeylanicus_populations_in_Singapore_a_last_straw_for_the_species

Long-term surveys reveal Singapore as the global stronghold of endangered songbird « Life Sciences « Cambridge Core Blog http://blog.journals.cambridge.org/2017/02/23/long-term-surveys-reveal-singapore-as-the-global-stronghold-of-endangered-songbird

The tiny corner of Asia where an Endangered songbird is thriving | @BirdLife_News

http://www.birdlife.org/worldwide/news/tiny-corner-asia-where-endangered-songbird-thriving

https://SingaporeBirdGroup.wordpress.com/2017/02/25/singapore-the-global-stronghold-of-the-straw-headed-bulbul/

---------------------------------------------------------------

روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

[Ornithology • 2019] Pycnonotus pseudosimplex • A Cryptic New Species of Bulbul (Pycnonotidae) from Borneo ---ScRaBBlE

Pycnonotus pseudosimplex

Shakya, Lim, Moyle, Rahman, Lakim & Sheldon, 2019

DOI 10.25226/bboc.v139i1.2019.a3

Abstract

Cream-vented Bulbul Pycnonotus simplex of Borneo was previously considered to be polymorphic in iris colour, having either red or white (creamy-yellow) irides. Mitochondrial DNA sequence comparisons, however, indicate that white- and red-eyed Bornean individuals are not closely related to one another. Instead, white-eyed birds are sister to Ashy-fronted Bulbul P. cinereifrons of Palawan Island, in the south-west Philippines, and red-eyed birds are sister to white-eyed P. simplex of the Thai-Malay Peninsula. Consequently, we elect to treat the white-eyed Bornean population as a distinct, previously overlooked species. In respect to plumage, white- and red-eyed individuals are almost identical, varying only slightly in the amount of yellow coloration in their feathers. The two taxa are sympatric at some localities, but white-eyed individuals are rarer and more consistently associated with mature forest than red-eyed birds.

Figure 3. Photographs from Lambir Hills National Park, Miri Division, Sarawak, Malaysia of
(A) Cream-eyed Bulbul Pycnonotus pseudosimplex sp. nov., and
(B) Cream-vented Bulbul P. simplex

Figure 2. Maximum likelihood tree generated from mitochondrial ND2 sequences. Numbers adjacent to branches indicate bootstrap support. Illustrations of heads of each species are shown to the right (depictions by Subir B. Shakya).

Taxonomy.— Genetic comparisons using mitochondrial ND2 sequences reveal that white- and red-eyed individuals of P. simplex from Borneo are not members of the same species. Their segregation by iris colour is backed by a deep ND2 sequence divergence (14%). Red-eyed Bornean birds are sister to white-eyed P. s. simplex of the Thai-Malay Peninsula at 4% ND2 divergence, whereas white-eyed Bornean birds are sister to P. cinereifrons of Palawan at 12% ND2 divergence. The perfect association of ND2 haplotype with iris colour across the region where the two morphotypes co-occur suggests assortative mating and the absence of mitochondrial gene introgression. For the red-eyed Bornean population, iris colour difference is adequate for recognition as a separate subspecies (P. s. perplexus; Chasen & Kloss, 1929, Eaton et al. 2016), or possibly even species, from white-eyed P. s. simplex of the Thai-Malay Peninsula and Sumatra. Because the two taxa are relatively closely related in respect to ND2 distance, species-level reclassification should depend on future research into gene flow and song. Plumage variation is unlikely to be helpful because of the similarity between these populations and the tendency for colours to change in museum specimens (Hoogerwerf 1966, Mees 1986). For the white-eyed Bornean population, its large genetic distance from P. cinereifrons and distinct morphology (Fig. 2) are sufficient to recognise it as a species.

There are no other taxa whose names apply to the Bornean white-eyed population. P. simplex subspecies have been described from Sumatra, the Anamba Islands, the northern Natuna Islands, the southern Natuna Islands, Java, and northern Borneo (Table 1, Fig. 1). Iris colour and size connect white-eyed populations of Sumatra and mainland Asia in subspecies simplex (Fig. 1). Larger size distinguishes the white-eyed subspecies halizonus (including its junior synonym axanthinus) of the Anamba and northern Natuna Islands from other white-eyed populations: e.g., mean wing length for halizonus 83.6 mm (n = 4) vs. 74 mm (n = 12) for Bornean white-eyed individuals. The red-eyed subspecies on the southern Natuna Islands (oblitus) belongs with Borneo's red-eyed population based on iris colour, size and plumage (Mees 1986). Eye colour of the Javan population (prillwitzi) varies from red to orange or red-brown, and its plumage is more distinctive than any other population of P. simplex, including red- and white-eyed Bornean populations (Hoogerwerf 1966). Apart from these taxa of P. simplex, there is no evidence that early taxonomists confused other species of similar-looking bulbuls with the white-eyed Bornean population of P. simplex. Pycnonotus brunneus has two subspecies: P. b. brunneus (including P. b. zaphaeus) and P. b. zapolius. Both of these were described as having red irides. P. erythropthalmos (including P. e. salvadorii, sometimes recognised for Borneo) has a red iris with a yellow eye-ring. Because no name applies to the Bornean white-eyed population, we name it:

Pycnonotus pseudosimplex, sp. nov.

Cream-eyed Bulbul

Diagnosis.—The iris of P. pseudosimplex is creamy yellow instead of crimson-red as in Cream-vented Bulbul P. s. simplex of Borneo. Its plumage also differs from that of P. s. simplex in being yellower on the throat and vent, creating greater contrast with the darker breast and flanks. From P. cinereifrons, P. pseudosimplex differs in being much smaller: P. cinereifrons mean 31.7 g (n = 3); P. pseudosimplex mean 20.7 g (n = 11). P. cinereifrons also has an olive tinge to the leading edge of its remiges, which is especially obvious on the folded wings of specimens (similar to P. plumosus of Borneo). From P. brunneus, P. pseudosimplex differs in having a white iris instead of an orange or two-toned iris. Similarly, P. erythropthalmos differs in having a dark red iris and a circle of yellow skin around the eyes.

Etymology.— For more than 100 years this species has been confused with P. simplex, i.e., since Carl Lumholtz collected the first white-eyed specimen in ‘Boeloengan’ (Balungan Regency) along the Kayan River in eastern Kalimantan, Indonesian Borneo, in 1914 (Lumholtz 1920, Voous 1961). Apart from eye colour, the two species are almost indistinguishable; hence, we refer to the new species as ‘false’ or ‘pseudo’ simplex. The common name, Cream-eyed Bulbul, describes the main identifying character of the new species. It also evokes a time when P. simplex was known as the White-eyed Brown Bulbul, as opposed to the Red-eyed Brown Bulbul P. brunneus of Borneo (Smythies 1960).

Habitat.— P. simplex and P. pseudosimplex appear to be sympatric in mature forest interior and edge. Individuals of both species whose DNA was compared in this study were collected in the same localities during recent years: viz., Ulu Kimanis, Crocker Range National Park, Sabah, in 2008; Lambir Hills National Park, Sarawak, in 2013; and Batang Ai National Park, Sarawak, in 2018 (Fig. 1, Table 2). Both taxa were also mist-netted together in the early 1980s elsewhere in Sabah: at Bole River, Saliwangan Baru, and Mangkatai (Megatai) (localities described in Sheldon 2015). Although previous authors have stated that white- and red-eyed individuals co-occur throughout Borneo (Hoogerwerf 1966, Mees 1986), our experience and specimen records suggest that white-eyed individuals are considerably rarer and perhaps more habitat-restricted. P. simplex occurs to 1,100 m (Kelabit Highlands) and in several forest types of varying quality: good-soil dipterocarp, kerangas and peatswamp; primary, secondary, and heavily disturbed forest; and plantations. We have found P. pseudosimplex (only by mist-netting) to 500 m (Ulu Kimanis) near the edge of mature good-soil dipterocarp, and in kerangas forest nearer to sea level.

Subir B. Shakya, Haw Chuan Lim, Robert G. Moyle, Mustafa Abdul Rahman, Maklarin Lakim and Frederick H. Sheldon. 2019. A Cryptic New Species of Bulbul from Borneo. Bulletin of the British Ornithologists’ Club. 139(1) DOI 10.25226/bboc.v139i1.2019.a3

---------------------------------------------------------------

روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

[Ornithology • 2018] Nok, gen. nov. • Phylogenetic Affinities of the Enigmatic Bare-faced Bulbul Pycnonotus hualon with Description of A New Genus ---ScRaBBlE

Nok hualon (Woxvold, Duckworth & Timmins, 2009)

in Fuchs, Pasquet, Stuart, Woxvold, Duckworth & Bowie, 2018.

DOI: 10.1111/ibi.12580

Abstract

Figure 1. Image of Bare-faced Bulbul “Pycnonotus hualon”. captured on the Pha Lom limestone massif, northeast Savannakhet Province, Lao PDR in December 2008.

(Photo: I. Woxvold).

by Risso Romain commons.wikimedia.org

Nok, gen. nov.

Type species: Pycnonotus hualon Woxvold, Duckworth and Timmins, 2009 by original description.

Included species: Nok hualon (Woxvold, Duckworth & Timmins, 2009), comb. nov.

ນົກຂວກຫົວລ້ານ / ນົກຂວາກຫົວລ້ານ

ResearchGate.net/publication/322697999_Phylogenetic_affinities_Pycnonotus_hualon

facebook.com/bcst.or.th/photos/1593960444052226

---------------------------------------------------------------

روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ScRaBBlE

Search This Blog

مشاركة مميزة

[Ornithology • 2017] Myzomela irianawidodoae • A Colourful New Species of Myzomela Honeyeater from Rote Island in eastern Indonesia ---ScRaBBlE

Translate

Wednesday, March 20, 2019

[Ornithology • 2018] Nok, gen. nov. • Phylogenetic Affinities of the Enigmatic Bare-faced Bulbul Pycnonotus hualon with Description of A New Genus ---ScRaBBlE

[Ornithology • 2017] Phylogeography of Bulbuls in the Genus Iole (Aves: Pycnonotidae) ---ScRaBBlE

[Ornithology / Conservation • 2017] Significance of the Globally Threatened Straw-headed Bulbul (Pycnonotus zeylanicus) Populations in Singapore: A Last Straw for the Species? ---ScRaBBlE

[Ornithology • 2019] Pycnonotus pseudosimplex • A Cryptic New Species of Bulbul (Pycnonotidae) from Borneo ---ScRaBBlE

[Ornithology • 2018] Nok, gen. nov. • Phylogenetic Affinities of the Enigmatic Bare-faced Bulbul Pycnonotus hualon with Description of A New Genus ---ScRaBBlE

المشاركات الشائعة