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[Ornithology • 2017] Myzomela irianawidodoae • A Colourful New Species of Myzomela Honeyeater from Rote Island in eastern Indonesia ---ScRaBBlE

Myzomela irianawidodoae Prawiradilaga, Baveja, Suparno, Ashari, Ng, Gwee, Verbelen & Rheindt, 2017  photo:   Philippe Verbelen  e-journ...

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Showing posts with label Saurischia. Show all posts
Showing posts with label Saurischia. Show all posts

Wednesday, March 20, 2019

[Paleontology • 2018] Sibirotitan astrosacralis • A New Sauropod dinosaur from the Lower Cretaceous Ilek Formation, Western Siberia, Russia ---ScRaBBlE


Sibirotitan astrosacralis
Averianov, Ivantsov, Skutschas, Faingertz & Leshchinskiy, 2018

Сибиротитан звездокрестцовый  DOI: 10.1016/j.geobios.2017.12.004

Abstract
Sibirotitan astrosacralis nov. gen., nov. sp., is described based on isolated but possibly associated cervical and dorsal vertebrae, sacrum, and previously published pedal elements from the Lower Cretaceous (Barremian?) Ilek Formation at Shestakovo 1 locality (Kemerovo Province, Western Siberia, Russia). Some isolated sauropod teeth from the Shestakovo 1 locality are referred to the same taxon. The phylogenetic parsimony analyses place Sibirotitan astrosacralis nov. gen., nov. sp., as a non-titanosaurian somphospondyl titanosauriform. The new taxon exhibits four titanosauriform and one somphospondylan synapomorphies, and one autapomorphy – a hyposphene ridge that extends between the neural canal and the postzygapophyses. It differs from all other Somphospondyli by having only five sacral vertebrae. The new taxon shares with Euhelopus and Epachtosaurus sacral ribs that converge towards the middle of the sacrum in dorsal view. Sibirotitan astrosacralis nov. gen., nov. sp., is only the second sauropod taxon from Russia and one of the oldest titanosauriform described so far in Asia.

Keywords: Sibirotitan astrosacralis nov. gen., nov. sp.; Titanosauriformes; Sauropoda; Phylogenetic parsimony analysis; Early Cretaceous; Siberia; Asia



Dinosauria Owen, 1842
Saurischia Seeley, 1887

Sauropoda Marsh, 1878
Titanosauriformes Salgado et al., 1997

Genus Sibirotitan nov. gen.

Derivation of the name: from Siberia and Greek Tιτάν (titan), a member of the second order of divine beings, descended from the primordial deities and preceding the Olympian deities in Greek mythology.

Type and only species: Sibirotitan astrosacralis nov. gen., nov. sp.

Derivation of the name: from Greek αστρον (star) and Latin os sacrum (“sacred bone”), an allusion to the unusual configuration of sacral ribs which radiate, in dorsal view, from the middle of the sacrum as the rays of a star.


 S.V. Ivantsov digging out the posterior cervical vertebra PM TGU 120/8-Sh1-3 from the Malyi Yar outcrop in 2008 (Kemerovo Province, Russia). Photo by S.V. Leshchinskiy.

  

  


 Alexander Averianov, Stepan Ivantsov, Pavel Skutschas, Alexey Faingertz and Sergey Leshchinskiy. 2018. A New Sauropod dinosaur from the Lower Cretaceous Ilek Formation, Western Siberia, Russia. Geobios. In Press.  DOI: 10.1016/j.geobios.2017.12.004



Averianov, A.O., Voronkevich, A.V., Maschenko, E.N., Leshchinskiy, S.V., and Fayngertz, A.V. 2002. A sauropod foot from the Early Cretaceous of Western Siberia, Russia. Acta Palaeontologica Polonica. 47 (1): 117–124.  DOI: 10.1.1.492.1575 


Познакомьтесь с новым сибирским динозавром https://metkere.com/2018/01/sibirotitan.html via @metkere
Новый род найденных в Кузбассе динозавров назвали сибиротитаном https://mediakuzbass.ru/news/obshhestvo/94522.html 


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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2018] A Revised Cranial Description of Massospondylus carinatus Owen (Dinosauria: Sauropodomorpha) based on Computed Tomographic Scans and A Review of Cranial Characters for Basal Sauropodomorpha ---ScRaBBlE


Massospondylus carinatus  Owen, 1854

in Chapelle & Choiniere, 2018
   DOI: 10.7717/peerj.4224 

Abstract

Massospondylus carinatus is a basal sauropodomorph dinosaur from the early Jurassic Elliot Formation of South Africa. It is one of the best-represented fossil dinosaur taxa, known from hundreds of specimens including at least 13 complete or nearly complete skulls. Surprisingly, the internal cranial anatomy of M. carinatus has never been described using computed tomography (CT) methods. Using CT scans and 3D digital representations, we digitally reconstruct the bones of the facial skeleton, braincase, and palate of a complete, undistorted cranium of M. carinatus (BP/1/5241). We describe the anatomical features of the cranial bones, and compare them to other closely related sauropodomorph taxa such as Plateosaurus erlenbergiensis, Lufengosaurus huenei, Sarahsaurus aurifontanalis and Efraasia minor. We identify a suite of character states of the skull and braincase for M. carinatus that sets it apart from other taxa, but these remain tentative due to the lack of comparative sauropodomorph braincase descriptions in the literature. Furthermore, we hypothesize 27 new cranial characters useful for determining relationships in non-sauropodan Sauropodomorpha, delete five pre-existing characters and revise the scores of several existing cranial characters to make more explicit homology statements. All the characters that we hypothesized or revised are illustrated. Using parsimony as an optimality criterion, we then test the relationships of M. carinatus (using BP/1/5241 as a specimen-level exemplar) in our revised phylogenetic data matrix.


Figure 1: Photographs of the skull of BP/1/5241. (A) Left lateral view. (B) Right lateral view. (C) Dorsal view.

Figure 2: Reconstructed skull of BP/1/5241.
(A) Left lateral view. (B) Right lateral view.

 aof, antorbital fenestra; aofs, antorbital fossa; bo, basioccipital; ecpt, ectopterygoid; eo, exoccipital; fr, frontal; itf, infratemporal fenestra; j, jugal; la, lacrimal; mx, maxilla; n, nasal; nf, narial fenestra; obt, orbit; pa, parietal; pf, prefrontal; pmx, premaxilla; po, postorbital; q, quadrate; qj, quadratojugal; sq, squamosal.

SYSTEMATIC PALEONTOLOGY
Dinosauria Owen, 1842
Saurischia Seeley, 1887
Sauropodomorpha Huene, 1932

Massospondylidae Huene, 1914 sensu Yates, 2003b

Massospondylus carinatus Owen, 1854

....

Conclusion: 
The use of CT scanning and 3D visualization graphics allows for a better understanding of the internal and external morphological structures of the braincase as well as information about the soft tissues such as the vestibular canals. M. carinatus can be tentatively diagnosed cranially by basipterygoid processes that are separated by an angle smaller than 60° and a jugal process of the ectopterygoid that is strongly curved. A revision of cranial characters provides a basis for more comparative work on the braincase of sauropodomorphs in general. Results also show a well-supported Massospondylidae clade. Further phylogenetic analyses using individual specimens of known ontogenetic stages as operational taxonomic units would provide better resolution for the Massospondylidae clade as well as a better understanding of which sets of character states set M. carinatus aside from other taxa.


Kimberley E.J. Chapelle and Jonah N. Choiniere. 2018. A Revised Cranial Description of Massospondylus carinatus Owen (Dinosauria: Sauropodomorpha) based on Computed Tomographic Scans and A Review of Cranial Characters for Basal Sauropodomorpha.   PeerJ. 6:e4224.  DOI: 10.7717/peerj.4224
CT-scan study makes it possible to 3-D print the skull of the dinosaur species Massospondylus http://phy.so/434970306 via @physorg_com



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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Mierasaurus bobyoungi • Descendants of the Jurassic Turiasaurs from Iberia Found Refuge in the Early Cretaceous of western USA ---ScRaBBlE


Mierasaurus bobyoungi 
Royo-Torres, Upchurch, Kirkland, DeBlieux, Foster, Cobos & Alcalá, 2017

reconstruction: Mike Skrepnick

Abstract
A new, largely complete eusauropod dinosaur with cranial and postcranial elements from two skeletons, Mierasaurus bobyoungi gen. nov., sp. nov. from the lower Yellow Cat Member (Early Cretaceous) of Utah (USA), is the first recognized member of Turiasauria from North America. Moreover, according to our phylogenetic results, Moabosaurus utahensis from the lower Yellow Cat Member of Utah (USA) is also a member of this clade. This group of non-neosauropod eusauropods, which now includes five genera (Losillasaurus, Turiasaurus, Mierasaurus, Moabosaurus and Zby), was previously known only from the Jurassic of Europe. These recent discoveries in Utah suggest that turiasaurs as a lineage survived the Jurassic-Cretaceous extinction boundary and expanded their known range, at least, into western North America. The revised spatiotemporal distribution of turiasaurs is consistent with the presence of a land connection between North America and Europe sometime during the late Tithonian to Valanginian (c.147-133 Ma). Mierasaurus and Moabosaurus are the only non-neosauropod eusauropods known from North America, despite being younger than the classic neosauropods of the Morrison Formation (c.150 Ma).






Figure 2: The skull material (UMNH.VP.26004) of Mierasaurus bobyoungi gen. nov, sp. nov.

Systematic Palaeontology

Dinosauria Owen, 1842
Saurischia Seeley, 1887

Sauropoda Marsh, 1878
Eusauropoda Upchurch, 1995
Turiasauria Royo-Torres, Cobos and Alcalá, 2006

Mierasaurus bobyoungi gen. nov., sp. nov.

Etymology: Genus named for Bernardo de Miera y Pacheco, Spanish cartographer and chief scientist for the 1776 Domínguez-Escalante Expedition: the first European scientist to enter what is now Utah. The species name acknowledges the importance of the underappreciated research by Robert Young on the Early Cretaceous of Utah.

Holotype: A partial skeleton of a single individual (UMNH.VP.26004), comprising disarticulated cranial and postcranial elements from the type site (Doelling’s Bowl). We regard this as subadult because it is a relatively large animal with unfused vertebral centra and neural arches in some dorsal vertebrae. This individual includes a partial skull and jaw, teeth, atlas, 8 cervical vertebrae, 11 cervical ribs, 11 dorsal vertebrae, 6 dorsal ribs, 6 sacral ribs, 15 caudal vertebrae, two chevrons, right scapula and partial left scapula, left radius, left ulna, left manus, complete pelvic elements, both femora, left tibia, left fibula, left astragalus and left pes.

tibia, fibula and complete left hind foot of Mierasaurus bobyoungi.



Type locality and horizon: All Mierasaurus remains discussed herein are from Doelling’s Bowl bonebed, UMNH VP.LOC.1208 (Utah Loc. 42Gr0300v) within the lower Yellow Cat Member (below the marker calcrete), Cedar Mountain Formation, on lands managed by the U.S. Bureau of Land Management in northern Grand County, east-central Utah. In a preliminary report in a conference abstract, detrital zircon dating indicates maximum ages ranging from ~136.4 ± 1.1 Ma and ~132 Ma for lower Yellow Cat Member and ~137.2 ± 2.0 Ma for upper Yellow Cat Member. The combination of these data with the upper Berriasian-Valanginian age based on ostracods and charophyte fauna for the upper Yellow Cat Member indicates a conflict in the maximum age traditionally considered for it 124.2 ± 2.6 Ma (see Supplementary Information). All of these data suggest a potential age of late Berriasian-early Aptian (c.142-124 Ma) for the Yellow Cat Member.

Doelling’s Bowl has produced the iguanodont cf. Iguanocolossus sp., a new species of polacanthid ankylosaur, a large allosauroid theropod (teeth), and the dromaeosaur Yurgovuchia doellingi. Exact locality information will be provided to qualified researchers on request through the Natural History Museum of Utah or the Utah Geological Survey.

Diagnosis: A turiasaurian sauropod possessing the following features (autapomorphies marked by *): *the otosphenoidal ridge extends from the anterior surface of the paroccipital process, near its ventral margin and is restricted to the medialmost part of the latter process (Fig. 2b); *the occipital condyle has a pair of rounded ridges extending dorsoventrally, one on either lateral face of the condylar articular surface (Fig. 2a–d); *the atlantal intercentrum (Fig. 3e,f) bears a pair of depressions in the medial surface, facing posteromedially, each of which receives the anterolateral margin of the odontoid process; a well-developed spinoprezygapophyseal lamina extends anteriorly (Fig. 3a–d, g–m) as a low ridge onto the lateral surface of the prezygapophyses roofing a lateral fossa on prezygapophyses in middle and posterior cervical vertebrae (shared with the diplodocine Kaatedocus26); *cervical ribs bear a ridge or bulge on the lateral surface of the tuberculum, immediately posterior to the base of the anterior process (Fig. 3n–v); dorsal neural arches lack posterior centroparapophyseal laminae; *lateral depression on the distal ramus of haemal arches (Fig. 5g,h); metacarpal I longer than metacarpal IV, shared with Macronaria27 (Fig. 5m); a very short ischium compared to pubis length (ischium:pubis length ratio = 0.75) (Fig. 5d–f); the midpoint of the fourth trochanter placed in the proximal part of the femur (Fig. 5a); femur with subequal distal condyles (Fig. 5k); and pedal unguals 2 and 3 compressed dorsoventrally (Fig. 5l,o).



Figure 6: Phylogenetic relationships of Turiasauria: The time-calibrated phylogenetic relationships of Mierasaurus bobyoungi n. gen. et sp. nov. The box next to each taxon demarcates its temporal range, whereas the colour of the box reflects the continent(s) where the taxon occurs (yellow = South America, light blue = Asia, orange = North America; green = Europe, dark blue = Africa; black = several continents).

 mired Mierasaurus type specimen.

reconstruction: Mike Skrepnick 






Rafael Royo-Torres, Paul Upchurch, James I. Kirkland, Donald D. DeBlieux, John R. Foster, Alberto Cobos & Luis Alcalá. 2017. Descendants of the Jurassic Turiasaurs from Iberia Found Refuge in the Early Cretaceous of western USA.  Scientific Reports. 7, Article number: 14311. DOI: 10.1038/s41598-017-14677-2

Identificado un nuevo #dinosaurio de #Utah (#EEUU) con orígenes en #Teruel (#España) 



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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Europatitan eastwoodi • A New Sauropod from the lower Cretaceous of Iberia in the Initial Radiation of Somphospondylans in Laurasia ---ScRaBBlE


Europatitan eastwoodi
Fernández-Baldor​, Canudo​, Pedro Huerta​, Moreno-Azanza​ & Montero​, 2017

.  DOI: 10.7717/peerj.3409 

Abstract

The sauropod of El Oterillo II is a specimen that was excavated from the Castrillo de la Reina Formation (Burgos, Spain), late Barremian–early Aptian, in the 2000s but initially remained undescribed. A tooth and elements of the axial skeleton, and the scapular and pelvic girdle, represent it. It is one of the most complete titanosauriform sauropods from the Early Cretaceous of Europe and presents an opportunity to deepen our understanding of the radiation of this clade in the Early Cretaceous and study the paleobiogeographical relationships of Iberia with Gondwana and with other parts of Laurasia. The late Barremian–early Aptian is the time interval in the Cretaceous with the greatest diversity of sauropod taxa described in Iberia: two titanosauriforms, Tastavinsaurus and Europatitan; and a rebbachisaurid, DemandasaurusThe new sauropod Europatitan eastwoodi n. gen. n. sp. presents a series of autapomorphic characters in the presacral vertebrae and scapula that distinguish it from the other sauropods of the Early Cretaceous of Iberia. Our phylogenetic study locates Europatitan as the basalmost member of the Somphospondyli, clearly differentiated from other clades such as Brachiosauridae and Titanosauria, and distantly related to the contemporaneous Tastavinsaurus. Europatitan could be a representative of a Eurogondwanan fauna like Demandasaurus, the other sauropod described from the Castrillo de la Reina Formation. The presence of a sauropod fauna with marked Gondwananan affinities in the Aptian of Iberia reinforces the idea of faunal exchanges between this continental masses during the Early Cretaceous. Further specimens and more detailed analysis are needed to elucidate if this Aptian fauna is caused by the presence of previously unnoticed Aptian land bridges, or it represents a relict fauna from an earlier dispersal event.

Figure 10: First caudal vertebra (MDS-OTII,2) of Europatitan eastwoodi n. gen. n. sp.
(A) Anterior view. (B) Right lateral view. (C) Posterior view. (D) Dorsal view. (E) Ventral view.

 ACDL, anterior centrodiapophyseal lamina; POSZG, postzygapophyses; PRDL, prezygodiapophyseal lamina; PRZG, prezygapophyses; SPOL, spinopostzygapophyseal lamina; SPRL, spinoprezygapophyseal lamina. Scale: 10 cm.

Figure 2: Quarry map of the partial skeleton of Europatitan eastwoodi n. gen. n. sp. from the late Barremian–early Aptian, Early Cretaceous, of El Oterillo II site, Spain.
The arrow indicates an iguanodontoid ilium (Contreras et al., 2007). Circular symbols correspond to splinters, and triangles to isolated teeth of theropods.


illustration: Davide Bonadona


Order SAURISCHIA Seeley, 1887
Infraorder SAUROPODA Marsh, 1878
NEOSAUROPODA Bonaparte, 1986

Titanosauriformes Salgado, Coria & Calvo, 1997
Somphospondyli Wilson & Sereno, 1998

Genus Europatitan gen. nov. 

Etymology: In reference to Europe, the continent where it was found, and the titans, ancient Greek deities known for their gigantic size, endowed with great power.

Type Species: Europatitan eastwoodi sp. nov. 

Etymology: Dedicated to US actor Clint Eastwood, the protagonist of the film “The Good, the Bad and the Ugly,” which was partially filmed near Salas de los Infantes.

Type Locality and Horizon: The site of El Oterillo II is located in the province of Burgos in northern Spain, 2.5 km to the west of the village of Barbadillo del Mercado in Salas de los Infantes (Fig. 1), Burgos Province, Spain; Urbión Group, Castrillo de la Reina Fm., lower Cretaceous, regarded as late Barremian–early Aptian in age (Martín-Closas & Alonso Millán, 1998).

Holotype: MDS-OTII,1 to MDS-OTII-32. The disarticulated carcass of a single specimen consisting of the following material: one tooth, five cervical vertebrae, one dorsal vertebra, nine caudal vertebrae, 11 cervical ribs, five dorsal ribs, seven hemal arches, the two scapulae, the left coracoid, the left metacarpals I and III, the two pubes, and the two ischia.

.....


Fidel Torcida Fernández-Baldor​, José Ignacio Canudo​, Pedro Huerta​, Miguel Moreno-Azanza​ and Diego Montero​. 2017. Europatitan eastwoodi, A New Sauropod from the lower Cretaceous of Iberia in the Initial Radiation of Somphospondylans in Laurasia.  
 PeerJ. 5:e3409.  DOI: 10.7717/peerj.3409

El Europatitan eastwoodi, un nuevo saurópodo hallado en la Sierra de La Demanda... http://burgosconecta.es/2017/06/28/el-europatitan-eastwoodi-el-nuevo-sauropodo-hallado-en-la-sierra-de-la-demanda/ via @burgosconecta  @gabi_iglesia  

       

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
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ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Jianianhualong tengi • Mosaic Evolution in An Asymmetrically Feathered Troodontid Dinosaur with Transitional Features ---ScRaBBlE


Jianianhualong tengi  
Xu, Currie, Pittman, Xing, Meng, Lü, Hu & Yu, 2017 

 Life reconstruction by Julius T. Csotonyi | DOI: 10.1038/ncomms14972


Abstract
Asymmetrical feathers have been associated with flight capability but are also found in species that do not fly, and their appearance was a major event in feather evolution. Among non-avialan theropods, they are only known in microraptorine dromaeosaurids. Here we report a new troodontid, Jianianhualong tengi gen. et sp. nov., from the Lower Cretaceous Jehol Group of China, that has anatomical features that are transitional between long-armed basal troodontids and derived short-armed ones, shedding new light on troodontid character evolution. It indicates that troodontid feathering is similar to Archaeopteryx in having large arm and leg feathers as well as frond-like tail feathering, confirming that these feathering characteristics were widely present among basal paravians. Most significantly, the taxon has the earliest known asymmetrical troodontid feathers, suggesting that feather asymmetry was ancestral to Paraves. This taxon also displays a mosaic distribution of characters like Sinusonasus, another troodontid with transitional anatomical features.

Figure 1: Jianianhualong tengi holotype DLXH 1218.
 (a) Photograph and (b) line drawing of the specimen. Scale bar, 50 cm.

 cav, caudal vertebrae; cv, cervical vertebrae; dv, dorsal vertebrae; fu, furcula; lfe, left femur; lh, left humerus; li, left ilium; lis, left ischium; lm, left manus; lp left pes; lpu, left pubis; lr, left radius; lu, left ulna; md, mandible; rc, right coracoid; rfe, right femur; rfi, right fibula; rh, right humerus; ri, right ilium; rm, right manus; rr, right radius; rs, right scapula; rt, right tibiotarsus; ru, right ulna; sk, skull; ss, synsacrum. 

Figure 7: Life reconstruction of Jianianhualong tengi based on the holotype DLXH 1218.
by Julius T. Csotonyi 

Systematic palaeontology 
Theropoda Marsh, 188121
Coelurosauria Huene, 192022

Maniraptora Gauthier, 198623
Troodontidae Gilmore, 192424

Jianianhualong tengi gen. et sp. nov.

 Etymology.Jianianhua’, the Chinese company that supported this study; ‘long’, the Chinese Pinyin for dragon. The specific name honors Ms Fangfang Teng, who secured the specimen for study.

 Holotype. DLXH 1218, a nearly complete skeleton with associated feathers (Fig. 1) housed at the Dalian Xinghai Museum.


Locality and horizon. Baicai Gou, Yixian County, western Liaoning, China; Lower Cretaceous Yixian Formation.

 Diagnosis. A troodontid distinguishable from other taxa in possessing the following apomorphic features (*indicates autapomorphic feature): maxillary rostral ramus triangular in outline and relatively high dorsoventrally*; maxillary ascending process extending posterodorsally at a high angle (an angle of ∼45° to maxillary ventral margin)*; lacrimal with a long descending process sub-equal in length to anterior process; a prominent ridge along anterior edge of the lateral surface of the lacrimal descending process; a distinct fossa on the dorsal surface of the surangular close to its posterior end; axial neural spine with a convex dorsal margin, transversely thickened anterior margin, and posterodorsal portion expanding strongly posteriorly; long manual phalanx II-1 (slightly shorter than metacarpal III) with prominent proximoventral heel, large groove along the medial surface of more than proximal half of manual phalanx II-1*; highly elongated manual III-2 (slightly longer than metacarpal III)*; robust ungual phalanges (medial ungual proximal depth/ungual length ratio >0.5); ilium with slightly concave dorsal margin in lateral view*; small medial lamina along ischial obturator process dorsal margin; metatarsal IV without prominent ventral flange*.

Figure 2: Cranium of Jianianhualong tengi holotype DLXH 1218.
 (a) Photograph and (b) line drawing of the skull and mandible. Scale bar, 2 cm.
ax, axis; fg, groove with foramina; fo, fossa; imf, internal mandibular fenestra; la, left angular; ld, left dentary; lect, left ectopterygoid; lex, left exoccipital; lf; left frontal; lhy, left hyoid bone; lj, left jugal; lla, left lacrimal; lm, left maxilla; lmf, left maxillary fenestra; ln, left nasal; lp, left parietal; lpt?, left pterygoid?; lpa, left prearticular; lpo, left postorbital; lpsaf, left posterior surangular foramen; lsa, left surangular; lsp?, left splenial?; lso, left supraoccipital; lsq, left squamosal; mc, Meckelian canal; ra, right angular; rart, right articular; rd, right dentary; rf, right frontal; rhy, right hyoid bone; rm?, right maxilla?; rn; right nasal; rp, right parietal; rpa, right prearticular; rq?, right quadrate?; rsa; right surangular; rso, right supraoccipital; rsp; right splenial; rsq, right squamosal. 



Xing Xu, Philip Currie, Michael Pittman, Lida Xing, Qingjin Meng, Junchang Lü, Dongyu Hu and Congyu Yu. 2017. Mosaic Evolution in An Asymmetrically Feathered Troodontid Dinosaur with Transitional Features.
  Nature Communications. 8, Article number: 14972.  DOI: 10.1038/ncomms14972

New Birdlike Dinosaur Had Modern Feathers on.natgeo.com/2p20KoN  @NatGeo

  


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او
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كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

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