[Paleontology • 2018] Osteology of the Late Cretaceous Argentinean Sauropod Dinosaur Mendozasaurus neguyelap: Implications for Basal Titanosaur Relationships ---ScRaBBlE

Mendozasaurus neguyelap Gonzàlez Riga, 2003

in Gonzàlez Riga, Mannion, Poropat, David & Coria. 2018. 

DOI: 10.1093/zoolinnean/zlx103  

Abstract

The titanosaurian sauropod dinosaur Mendozasaurus neguyelap is represented by several partial skeletons from a single locality within the Coniacian (lower Upper Cretaceous) Sierra Barrosa Formation in the south of Mendoza Province, northern Neuquén Basin, Argentina. A detailed revision of Mendozasaurus, including previously undocumented remains from the holotype site, allows us to more firmly establish its position within Titanosauria, as well as enabling an emended diagnosis of this taxon. Autapomorphies include: (1) middle and posterior cervical vertebrae with tall and transversely expanded neural spines that are wider than the centra, formed laterally by spinodiapophyseal laminae that are not connected with the pre- or postzygapophyses; (2) anterior caudal vertebrae (excluding anteriormost) with ventrolateral ridge-like expansion of prezygapophyses; and (3) humerus with divided lateral distal condyle on anterior surface. New remains demonstrate that the presacral vertebrae of Mendozasaurus were not unusually short anteroposteriorly, with this compression instead resulting from taphonomic crushing. Comparative studies of articulated pedes of other taxa allow us to interpret that the pedal formula of Mendozasaurus was 2-2-2-2-0, based on disarticulated bones that form a right hind foot. Mendozasaurus was incorporated into an expanded version of a titanosauriform-focussed phylogenetic data matrix, along with several other contemporaneous South American titanosaurs. The resultant data matrix comprises 84 taxa scored for 423 characters, and our phylogenetic analysis recovers Mendozasaurus as the most basal member of a diverse Lognkosauria, including Futalognkosaurus and the gigantic titanosaurs Argentinosaurus, Notocolossus, Patagotitan and Puertasaurus. Lognkosauria forms a clade with Rinconsauria (Muyelensaurus + Rinconsaurus), with Epachthosaurus and Pitekunsaurus recovered at the base of this grouping. A basal lithostrotian position for this South American clade is well supported, contrasting with some analyses that have placed these taxa outside of Lithostrotia or closer to Saltasauridae. The sister clade to this South American group is composed of an array of near-global taxa and supports the hypothesis that most titanosaurian clades were widespread by the Early–middle Cretaceous.

Figure 3. Life restoration of Mendozasaurus neguyelap based on the largest adult individual (femur length: 1530 mm). Artwork by Bernardo Gonzaléz Riga.

  CONCLUSIONS

A detailed description of all remains pertaining to the early Late Cretaceous Argentinean titanosaurian sauropod dinosaur Mendozasaurus neguyelap enables a revised diagnosis for the genus. An expanded phylogenetic analysis recovers Mendozasaurus and several other taxa as part of a rich Lognkosauria that is placed within a diverse clade of South American lithostrotian titanosaurs. The sister clade to this South American group is composed of a near-global array of titanosaurs, which supports recent work that has argued for a widespread distribution of most titanosaurian clades by the Early–middle Cretaceous.

Bernardo J. Gonzàlez Riga, Philip D. Mannion, Stephen F. Poropat, Leonardo D. Ortiz David and  Juan Pedro Coria. 2018. Osteology of the Late Cretaceous Argentinean Sauropod Dinosaur Mendozasaurus neguyelap: Implications for Basal Titanosaur Relationships. Zoological Journal of the Linnean Society. zlx103.  DOI: 10.1093/zoolinnean/zlx103

 facebook.com/photo.php?fbid=10210673274980433

Bernardo J. González Riga. 2003. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza, Argentina. Amehginiana. 40: 155-172.

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] A New, Exceptionally Preserved Juvenile Specimen of Eusaurosphargis dalsassoi (Diapsida) and Implications for Mesozoic Marine Diapsid Phylogeny ---ScRaBBlE

Eusaurosphargis dalsassoi Nosotti & Rieppel, 2003

Illustration: Beat Scheffold
DOI: 10.1038/s41598-017-04514-x 

Abstract

Recently it was suggested that the phylogenetic clustering of Mesozoic marine reptile lineages, such as thalattosaurs, the very successful fish-shaped ichthyosaurs and sauropterygians (including plesiosaurs), among others, in a so-called ‘superclade’ is an artefact linked to convergent evolution of morphological characters associated with a shared marine lifestyle. Accordingly, partial ‘un-scoring’ of the problematic phylogenetic characters was proposed. Here we report a new, exceptionally preserved and mostly articulated juvenile skeleton of the diapsid reptile, Eusaurosphargis dalsassoi, a species previously recovered within the marine reptile ‘superclade’, for which we now provide a revised diagnosis. Using micro-computed tomography, we show that besides having a deep skull with a short and broad rostrum, the most outstanding feature of the new specimen is extensive, complex body armour, mostly preserved in situ, along its vertebrae, ribs, and forelimbs, as well as a row of flat, keeled ventrolateral osteoderms associated with the gastralia. As a whole, the anatomical features support an essentially terrestrial lifestyle of the animal. A review of the proposed partial character ‘un-scoring’ using three published data matrices indicate that this approach is flawed and should be avoided, and that within the marine reptile ‘superclade’ E. dalsassoi potentially is the sister taxon of Sauropterygia.

A life reconstruction of Eusaurosphargis dalsassoi based on new specimen PIMUZ A/III 4380.

Illustration: Beat Scheffold. 

Figure 2: Eusaurosphargis dalsassoi (PIMUZ A/III 4380) in ventral view. (a) Photograph. (b) Interpretative drawing of skeleton. Abbreviations used in the figure are chevron (ch), gastralia (g), and lateral osteoderm (lo). 

Systematic Palaeontology

Diapsida Osborn, 190321.

Eusaurosphargis dalsassoi Nosotti and Rieppel, 2003.

Holotype: BES SC 390 (Palaeontological Collection of the Museo Civico di Storia Naturale di Milano, Italy), an associated but disarticulated specimen from the Besano Formation (242 Ma, Anisian-Ladinian, Middle Triassic) of Besano, Lombardy, Italy.

Referred specimen: PIMUZ A/III 4380, an almost completely articulated skeleton preserved in ventral view (Fig. 2a).

Stratigraphy and locality: Upper Prosanto Formation (bed no. 1; fossil found on bottom side of a limestone slab); 241 Ma, early Ladinian, Middle Triassic of Ducanfurgga locality no. 4, south of Davos, Canton Grisons (Graubünden), south-eastern Switzerland.

  

Torsten M. Scheyer, James M. Neenan, Timea Bodogan, Heinz Furrer, Christian Obrist and Mathieu Plamondon. 2017. A New, Exceptionally Preserved Juvenile Specimen of Eusaurosphargis dalsassoi (Diapsida) and Implications for Mesozoic Marine Diapsid Phylogeny.
 Scientific Reports. 7, 4406.  DOI: 10.1038/s41598-017-04514-x

Rare, exceptionally preserved fossil reveals lifestyle of ancient armor-plated reptile  phy.so/418006972   @physorg_com

Davoser Saurier zeigt erstmals seine bizarre Schuppenpanzerung
 http://www.idw-online.de/-ClXPAA

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2018] Osteology of the Late Cretaceous Argentinean Sauropod Dinosaur Mendozasaurus neguyelap: Implications for Basal Titanosaur Relationships ---ScRaBBlE

Mendozasaurus neguyelap Gonzàlez Riga, 2003

in Gonzàlez Riga, Mannion, Poropat, David & Coria. 2018. 

DOI: 10.1093/zoolinnean/zlx103  

Abstract

The titanosaurian sauropod dinosaur Mendozasaurus neguyelap is represented by several partial skeletons from a single locality within the Coniacian (lower Upper Cretaceous) Sierra Barrosa Formation in the south of Mendoza Province, northern Neuquén Basin, Argentina. A detailed revision of Mendozasaurus, including previously undocumented remains from the holotype site, allows us to more firmly establish its position within Titanosauria, as well as enabling an emended diagnosis of this taxon. Autapomorphies include: (1) middle and posterior cervical vertebrae with tall and transversely expanded neural spines that are wider than the centra, formed laterally by spinodiapophyseal laminae that are not connected with the pre- or postzygapophyses; (2) anterior caudal vertebrae (excluding anteriormost) with ventrolateral ridge-like expansion of prezygapophyses; and (3) humerus with divided lateral distal condyle on anterior surface. New remains demonstrate that the presacral vertebrae of Mendozasaurus were not unusually short anteroposteriorly, with this compression instead resulting from taphonomic crushing. Comparative studies of articulated pedes of other taxa allow us to interpret that the pedal formula of Mendozasaurus was 2-2-2-2-0, based on disarticulated bones that form a right hind foot. Mendozasaurus was incorporated into an expanded version of a titanosauriform-focussed phylogenetic data matrix, along with several other contemporaneous South American titanosaurs. The resultant data matrix comprises 84 taxa scored for 423 characters, and our phylogenetic analysis recovers Mendozasaurus as the most basal member of a diverse Lognkosauria, including Futalognkosaurus and the gigantic titanosaurs Argentinosaurus, Notocolossus, Patagotitan and Puertasaurus. Lognkosauria forms a clade with Rinconsauria (Muyelensaurus + Rinconsaurus), with Epachthosaurus and Pitekunsaurus recovered at the base of this grouping. A basal lithostrotian position for this South American clade is well supported, contrasting with some analyses that have placed these taxa outside of Lithostrotia or closer to Saltasauridae. The sister clade to this South American group is composed of an array of near-global taxa and supports the hypothesis that most titanosaurian clades were widespread by the Early–middle Cretaceous.

Figure 3. Life restoration of Mendozasaurus neguyelap based on the largest adult individual (femur length: 1530 mm). Artwork by Bernardo Gonzaléz Riga.

  CONCLUSIONS

A detailed description of all remains pertaining to the early Late Cretaceous Argentinean titanosaurian sauropod dinosaur Mendozasaurus neguyelap enables a revised diagnosis for the genus. An expanded phylogenetic analysis recovers Mendozasaurus and several other taxa as part of a rich Lognkosauria that is placed within a diverse clade of South American lithostrotian titanosaurs. The sister clade to this South American group is composed of a near-global array of titanosaurs, which supports recent work that has argued for a widespread distribution of most titanosaurian clades by the Early–middle Cretaceous.

Bernardo J. Gonzàlez Riga, Philip D. Mannion, Stephen F. Poropat, Leonardo D. Ortiz David and  Juan Pedro Coria. 2018. Osteology of the Late Cretaceous Argentinean Sauropod Dinosaur Mendozasaurus neguyelap: Implications for Basal Titanosaur Relationships. Zoological Journal of the Linnean Society. zlx103.  DOI: 10.1093/zoolinnean/zlx103

 facebook.com/photo.php?fbid=10210673274980433

Bernardo J. González Riga. 2003. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza, Argentina. Amehginiana. 40: 155-172.

---------------------------------------------------------------

روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.