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[Ornithology • 2017] Myzomela irianawidodoae • A Colourful New Species of Myzomela Honeyeater from Rote Island in eastern Indonesia ---ScRaBBlE

Myzomela irianawidodoae Prawiradilaga, Baveja, Suparno, Ashari, Ng, Gwee, Verbelen & Rheindt, 2017  photo:   Philippe Verbelen  e-journ...

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Showing posts with label Thailand. Show all posts
Showing posts with label Thailand. Show all posts

Wednesday, March 20, 2019

[Botany • 2018] Thismia thaithongiana • A New Species of Mycoheterotroph (Dioscoreaceae: Thismieae) from An Unusual Habitat in Thailand ---ScRaBBlE


Thismia thaithongiana
 Chantanaorr. & Suddee

in Chantanaorrapint & Suddee, 2018.

With more than 60 currently accepted species, Thismia Griffith (1844: 221) is the largest genus of the tribe Thismieae of Dioscoreaceae (sensu APG 2016, or Thismiaceae of other authors). The genus is widely distributed mainly in the tropical and subtropical regions with a concentration of species in Southeast Asia (ca. 30 species) including the Malay Peninsula, Sumatra, Java, Borneo, Thailand and Vietnam. In last decade, many new taxa have been described from Southeast Asia (e.g. Larsen & Averyanov 2007, Chantanaorrapint 2008 2012, Tsukaya & Okada 2012, Dančák et al. 2013, Nuraliev et al. 2014 2015, Truong et al. 2014, Tsukaya et al. 2014, Chantanaorrapint & Sridith 2015, Hroneš et al. 2015, Chantanaorrapint et al. 2016, Sochor et al. 2017). Members of the genus are small mycoheterotrophic herbs with a highly reduced habit and usually grow among leaf litter in shady wet forests.

Keywords: Doi Hua Mot; Umphang; mycoheterotrophic; taxonomy; Thailand; Thismia; Monocots





พิศวงตานกฮูกหรือพิศวงไทยทอง Thismia thaithongiana Chantanaorr. & Suddee ตีพิมพ์ในวารสาร Phytotaxa. 333(2): 287–292. 2018. คำระบุชนิดตั้งให้เป็นเกียรติแก่ รศ.ดร.อบฉันท์ ไทยทอง


 Sahut Chantanaorrapint and Somran Suddee. 2018. Thismia thaithongiana (Dioscoreaceae: Thismieae), A New Species of Mycoheterotroph from An Unusual Habitat. Phytotaxa. 333(2); 287–292. DOI:  10.11646/phytotaxa.333.2.14


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او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


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تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Botany • 2017] A Revision of Microchirita (Gesneriaceae) in Thailand ---ScRaBBlE



 Puglisi & Middleton, 2017. 
Gardens' Bulletin Singapore. 69(2)

ABSTRACT

 Microchirita (C.B.Clarke) Yin Z.Wang (Gesneriaceae: Didymocarpoideae) in Thailand is revised and 29 species are recognised, two of which have three varieties each. Eight new species are described, Microchirita albocyanea C.Puglisi, Microchirita glandulosa C.Puglisi, Microchirita hypocrateriformis C.Puglisi, Microchirita limbata C.Puglisi, Microchirita luteola C.Puglisi, Microchirita tadphoensis C.Puglisi, Microchirita tetsanae C.Puglisi, Microchirita thailandica C.Puglisi; three new varieties are described, Microchirita involucrata var. gigantiflora C.Puglisi, Microchirita mollissima var. glabra C.Puglisi, Microchirita mollissima var. glandulophylla C.Puglisi; and one name is combined at a new rank, Microchirita involucrata var. capitis (Craib) C.Puglisi. Two lectotypifications are made, one of which is a second step lectotypification. A key to all taxa is given, all taxa are described, and many are illustrated. 

Keywords. Chirita, Didymocarpoideae, Flora of Thailand, Gesneriads, new species, taxonomy


Fig. 1. Inflorescence types.
A. Cristate inflorescence of Microchirita bimaculata (D.Wood) A.Weber & D.J.Middleton. From Middleton, D.J. et al 4514. B. Bracteate inflorescence of Microchirita rupestris (Ridl.) A.Weber & Rafidah. From Puglisi, C. et al. CP409. (Photos: A, D.J. Middleton; B, P. Karaket)

Taxonomic treatment 

Microchirita (C.B.Clarke) Yin Z.Wang,
J. Syst. Evol. 49: 59 (2011). 
– Chirita sect. Microchirita C.B.Clarke in Candolle & Candolle, Monogr. Phan. 5: 127 (1883).
 – Roettlera sect. Microchirita (C.B.Clarke) Fritsch in Engler & Prantl, Nat. Pflanzenfam. IV/3b: 148 (1895).
 – Didymocarpus sect. Microchirita (C.B.Clarke) Chun, Sunyatsenia 6: 290 (1946). 

– TYPE: Microchirita hamosa (R.Br.) Yin Z.Wang, lectotype designated by Burtt (1954: 196).



1. Microchirita albiflora D.J.Middleton & Triboun, Thai Forest Bull., Bot. 41: 19 (2013). 
– TYPE: Thailand, Chiang Rai, Mae Fa Luang District, ..., 1000 m alt., 23 September 2008, Middleton, D.J., Karaket, P., Triboun, P., Kawatkul, U. & Meeboonya, R. 4567 (holotype BKF; isotypes BK, E [E00629491], K, P [P00966762], QBG, SING [SING0229831]).  


2. Microchirita albocyanea C.Puglisi, sp. nov. 
Most similar to Microchirita limbata C.Puglisi in the overall shape of the corolla and in colour, but differs in not having a glandular indumentum and in the much longer corolla and larger calyx. – TYPE: Thailand, Loei, Pha Khao, ..., 447 m, 5 November 2014, Tetsana, N. et al. 876 (holotype BKF; isotype SING). (Fig. 2D–F)


3. Microchirita aratriformis (D.Wood) A.Weber & D.J.Middleton, Taxon 60: 778 (2011).
 – Chirita aratriformis D.Wood, Notes Roy. Bot. Gard. Edinburgh 31: 367 (1972); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 197 (1974); Burtt, Thai Forest Bull., Bot. 29: 87 (2001).
 – TYPE: [Vietnam], Tonkin, Langson, Khanmoi, Eberhardt 3332 (holotype P [P00602506]). (Fig. 3A–B)


4. Microchirita bimaculata (D.Wood) A.Weber & D.J.Middleton, Taxon 60: 778 (2011).
Chirita bimaculata D.Wood, Notes Roy. Bot. Gard. Edinburgh 31: 368 (1972); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 196 (1974); Burtt, Thai Forest Bull., Bot. 29: 87 (2001).
 – TYPE: Thailand, Maeklang Falls, c. 50 km Northwest of Chiang Mai, c. 430 m, 3 November 1967, Burtt, B.L. 5611 (holotype E [E00155280]). (Fig. 4)



Fig. 2.  ข้าวตอกโยนก Microchirita albiflora D.J.Middleton & Triboun. A. Habit. B. Front view of flower.All from Middleton, D.J. et al. 4567. Microchirita albocyanea C.Puglisi. E. Side view of the flower. F. Front view of the flower. All from Tetsana, N. et al. 876. (Photos: A, B, P. Karaket; C, D.J. Middleton; D–F, N. Tetsana)

Fig. 3. Microchirita aratriformis (D.Wood) A.Weber & D.J.Middleton. A. Side view of the flower. B. Front view of flower. All from Tetsana, N. et al. 871. Microchirita hamosa (R.Br.) Yin Z.Wang.   E. Side view of the flower. F. Front view of flower. C from Middleton, D.J. et al. 4519; D from Middleton, D.J. et al. 4522; E, F from Middleton, D.J. et al. 5016. (Photos: A, B, N. Tetsana; C, D, D.J. Middleton; E, F, P. Karaket)

Fig. 4. Microchirita bimaculata (D.Wood) A.Weber & D.J.Middleton.C. Detail of a unifoliate plant. D. Front view of the flower.  A, C from Suddee, S. et al. 4970; B from Middleton, D.J. et al. 4514; D from Middleton, D.J. et al. 4520; E, F from Middleton, D.J. et al. 4479. (Photos: A, C, S. Suddee; B, D–F, P. Karaket)

Fig. 5. Microchirita hemratii C.Puglisi. A. Front view of flower. B. Side view of the flower. All from Middleton, D.J. et al. 5775. Microchirita huppatatensis C.Puglisi. C. Detail of the flower. D. Habit. All from Middleton, D.J. et al. 5689. (Photos: P. Karaket

Fig. 6. Microchirita hypocrateriformis C.Puglisi.  C. Front view of a white flower. D. Side view of the flower. E. Front view of a blue flower. F. Blue-flowered caulescent plant. A–D from Tetsana, N. et al. 888; E, F from Tetsana, N. et al. 834. (Photos: N. Tetsana

Fig. 7. Microchirita involucrata (Craib) Yin Z.Wang var. involucrata. A. Habit and fruit. B. Front view of the flower.  All from Middleton, D.J. et al. 5391. (Photos: P. Karaket)

5. Microchirita elphinstonia (Craib) A.Weber & D.J.Middleton, Taxon 60: 778 (2011).
Chirita elphinstonia Craib, Bull. Misc. Inform. Kew 149 (1932); Barnett, Fl. Siam. 3: 224 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 195 (1974); Burtt, Thai Forest Bull., Bot. 29: 87 (2001). 

– TYPE: Cult. in Hort. Aberdeen from seeds of Marcan 2561, coll. Thailand, Krabin, Ban Keng, 30 m, limestone hill (lectotype K [K000545615], ...


6. Microchirita glandulosa C.Puglisi, sp. nov.
 Similar to Microchirita involucrata (Craib) Yin Z.Wang and M. rupestris (Ridl.) A.Weber & Rafidah) in having bracteate inflorescences. Differs from both in the bracts being fused only at the base (i.e. not divided as in Microchirita involucrata and not fused into a cup as in M. rupestris), in the dimorphic indumentum of sparse, long eglandular hairs and dense short glandular hairs on the leaf (eglandular indumentum in M. involucrata and M. rupestris), and in the tripartite calyx. It differs further from Microchirita involucrata in the serrate margin of the bracts and from M. rupestris in the much smaller size of the bracts. – TYPE: Thailand, Nan, Song Kwaw, Sakoen, Khao Tham Plakang, 750 m, 3 September 2006, Watthana, S. 2126 (holotype QBG; isotype CMU).



7. Microchirita hamosa (R.Br.) Yin Z.Wang, J. Syst. Evol. 49: 60 (2011).
 – Chirita hamosa R.Br., Cyrtandreae 117 (1839); Barnett, Fl. Siam. 3: 224 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 191 (1974); Burtt, Thai Forest Bull., Bot. 29: 87 (2001). 

– TYPE (conserved – see Middleton & Puglisi, 2015): Thailand, Tak, Umphang, ..., 915 m, 17 October 2014, Middleton, D.J., Hemrat, C., Karaket, P., Puglisi, C. & Suddee, S. 5762 (holotype E; isotypes BKF, SING). (Fig. 3C–F)


8. Microchirita hemratii C.Puglisi, Kew Bull. 71(1)-2: 4 (2016). 
– TYPE: Thailand, Tak, Mae Sot distr., Wat Tham Inthanin, 660 m, 18 October 2014, Middleton, D.J., Hemrat, C., Karaket, P., Puglisi, C. & Suddee S. 5775 (holotype BKF; isotypes E [E00663027], SING). (Fig. 5A–B)

9. Microchirita huppatatensis C.Puglisi, Kew Bull. 71(1)-2: 2 (2016). 
– TYPE: Thailand, Uthai Thani, Lan Sak, Huppatat Non Hunting Area, 122 m, 14 October 2014, Middleton, D.J., Hemrat, C., Karaket, P., Puglisi, C., Suddee, S. 5689 (holotype BKF). (Fig. 5C–D)


  


หยาดสวนสวรรค์ Microchirita hypocrateriformis C. Puglisi 
  ไม้ล้มลุกขึ้นตามเขาหินปูน ลำต้นอวบน้ำ ดอกสีเหลืองอ่อนหรือสีม่วง แต่ที่พบมากจะเป็นสีเหลืองอ่อน กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก 
พรรณไม้ถิ่นเดียวของไทย (endemic) พบขึ้นตามเขาหินปูนในภาคตะวันออกเฉียงเหนือของไทย เช่นบริเวณสวนสวรรค์ สวนหินผางาม อำเภอหนองหิน จังหวัดเลย อำเภอคอนสาร จังหวัดชัยภูมิ



10. Microchirita hypocrateriformis C.Puglisi, sp. nov. Differs from all other species of Microchirita (C.B.Clarke) Yin Z.Wang in the combination of long, narrow corolla tube, abruptly opening into spreading limb, in the long lower corolla lobe, and in the presence of a fringe of glandular indumentum at the base of the upper lip. 

– TYPE: Thailand, Chaiyaphum, Khon Sarn, Wat Tham Huang Po, 400 m, 19 October 2015, Suddee, S., Keiwbang, W., Hemrat, C. 4967 (holotype BKF; isotype SING). (Fig. 6)


11. Microchirita involucrata (Craib) Yin Z.Wang, J. Syst. Evol. 49: 60 (2011); Rafidah, Gard. Bull. Singapore 69: 15 (2017). 
– Chirita involucrata Craib, Gard. Chron., Ser. 3, 83: 140 (1928); Barnett, Fl. Siam. 3: 223 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 199 (1974); Burtt, Thai Forest Bull., Bot. 29: 88 (2001). 

– TYPE: Cult. Hort. Bot. Aberdeen from seeds collected by Kerr (Kerr 11172), Thailand, Kaw Tao [Surat Thani, Kao Tao], 30/12/1926 (lectotype ABD [specimen with appended protologue], designated by Puglisi in Rafidah (2017); isolectotypes ABD [2 sheets]). (Fig. 7)

11a. Microchirita involucrata var. involucrata 
11b. Microchirita involucrata var. capitis (Craib) C.Puglisi, stat. nov.
Chirita capitis Craib, Bull. Misc. Inform. Kew 173 (1930); Barnett, Fl. Siam. 3: 223 (1962).

11c. Microchirita involucrata var. gigantiflora C.Puglisi, var. nov. 

12. Microchirita karaketii D.J.Middleton & Triboun, Thai Forest Bull., Bot. 41: 17 (2013). 
– TYPE: Thailand, Chiang Mai, Chiang Dao District, Doi Chiang Dao Wildlife Sanctuary, Tam Pak Piang, 530 m alt., 20 September 2008, Middleton, D.J., Karaket, P., Triboun, P., Kawatkul, U. & Meeboonya, R. 4526 (holotype BKF; isotypes E [E00629480], P [P00966764], QBG). (Fig. 8) 



13. Microchirita lilacina C.Puglisi, Kew Bull. 71(1)-2: 5 (2016).
 – TYPE: Thailand, Tak, Umphang, 504 m, 15 October 2014, Middleton, D.J., Hemrat, C., Karaket, P., Puglisi, C. & Suddee, S. 5704 (holotype BKF; isotypes AAU, E [E00663028], K, QBG, SING). (Fig. 9)


หยาดผาผึ้ง Microchirita limbata C. Puglisi (Gesneriaceae) 
จากเขตรักษาพันธุ์สัตว์ป่าผาผึ้ง อำเภอคอนสาร จังหวัดชัยภูมิ ไม้ล้มลุกขึ้นตามก้อนหินปูน ลำต้นอวบน้ำ ดอกสีม่วง กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก 
เป็นพรรณไม้ชนิดใหม่ของโลกและเป็นพรรณไม้ถิ่นเดียวของไทย (endemic) ตีพิมพ์ในวารสาร Gardens' Bulletin Singapore 69(2): 211–284. ตัวอย่างต้นแบบ Suddee, Keiwbang & Hemrat 4968 เก็บจากเขาหินปูน ถ้ำฮวงโป เขตรักษาพันธุ์สัตว์ป่าผาผึ้ง 


14. Microchirita limbata C.Puglisi, sp. nov. 
Species characterised by the tubular corolla with white tube and blue lobes, and by the widespread glandular indumentum. It is most similar to Microchirita albocyanea C.Puglisi in the overall shape and colour of the corolla, but differs in the smaller flowers and in having a glandular indumentum. 

– TYPE: Thailand, Chaiyaphum, Khon San, Wat Tham Huang Po, 443 m, 19 October 2015, Suddee, S., Keiwbang, W., Hemrat, C. 4968 (holotype BKF; isotype SING). (Fig. 10)


Fig. 7. Microchirita involucrata (Craib) Yin Z.Wang var. involucrata.  C. Side view of the flower. D. Lateral view. All from Middleton, D.J. et al. 5391. (Photos: P. Karaket)
Fig. 8. Microchirita karaketii D.J.Middleton & Triboun.   C. Side view of the flower. D. Front view of the flower. A, C, D from Middleton, D.J. et al. 4526; B from Middleton, D.J. et al. 4536. (Photos: A–C, D.J. Middleton; D, P. Karaket)
Fig. 9. Microchirita lilacina C.Puglisi.  C. Cristate inflorescence, fruit and side view of the flower. D. Front view of the flower. A from Middleton, D.J. et al. 5704; B–D from Middleton, D.J. et al. 5699. (Photos: P. Karaket
Fig. 10. Microchirita limbata C.Puglisi. A. Habit. B. Front view of the flower. C. Fruits. D. Lateral view of the flower. A, D from Tetsana, N. et al. 883; B, C from Suddee, S. et al. 4968. (Photos: A, D, N. Tetsana; B, C, S. Suddee


Fig. 11. Microchirita luteola C.Puglisi. A. Habit. B. Front view of the flower.  All from Tetsana, N. et al. 829. Microchirita marcanii (Craib) A.Weber & D.J.Middleton.   E. Side view of the flower. F. Front view of the flower. All from RBG Edinburgh accession number 20121420. (Photos: A–C, N. Tetsana; D–F, D.J. Middleton)

Fig. 13.  Microchirita purpurea D.J.Middleton & Triboun. D. Habit. E. Lateral view of the flower. F. Front view of the flower. All from Middleton, D.J. et al. 5681. (Photos: P. Karaket
Fig. 14. Microchirita rupestris (Ridl.) A.Weber & Rafidah.   C. Side view of the flower. D. Detail of the corolla. A, B from Middleton, D.J. et al. 5721; C from Middleton, D.J. et al. 4836; D from Middleton, D.J. et al. 5204. (Photos: A, B, P. Karaket; C, D, D.J. Middleton
Fig. 18. Microchirita viola (Ridl.) A.Weber & Rafidah. A. Lateral view of the flower. B. Front view of the flower. All from Middleton, D.J. et al. 5467.   (Photos: A, B, D.J. Middleton)  


15. Microchirita luteola C.Puglisi, sp. nov. 
Similar to Microchirita tubulosa (Craib) A.Weber & D.J.Middleton but differs in not having spots inside the lateral corolla lobes, having an entire disk (usually dorsally cleft in M. tubulosa), glandular indumentum on the stems (vs. eglandular), and acuminate calyx lobes (vs. usually acute, more rarely slightly acuminate). It is also similar to Microchirita marcanii in the shape of the corolla, but differs in the mixed eglandular and glandular indumentum on many plant parts (eglandular only in M. marcanii (Craib) A.Weber & D.J.Middleton) and the corolla colour pattern (light yellow corolla with a yellow stripe vs. orange corolla with lateral purple spots). Finally, it differs from Microchirita elphinstonia (Craib) A.Weber & D.J.Middleton in having a glandular indumentum and in the larger and much paler yellow corolla. 
– TYPE: Loei, Nong Hin, Suan Sa Wan, Pha Ngam Forest Park, 662 m, 11 September 2014, Tetsana, N. et al. 829 (holotype BKF; isotype SING). (Fig. 11A–C)

16. Microchirita marcanii (Craib) A.Weber & D.J.Middleton, Taxon 60: 778 (2011). 
Chirita marcanii Craib, Bull. Misc. Inform. Kew 171 (1926); Barnett, Fl. Siam. 3: 226 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 193 (1974); Burtt, Thai Forest Bull., Bot. 29: 88 (2001). 
– TYPE: Thailand, Saraburi, Muak Lek, c. 250 m, 10 November 1924, Marcan 1872 (lectotype ABD, designated by Wood (1974: 193); isolectotypes K (2)). (Fig. 11D–F)


17. Microchirita micromusa (B.L.Burtt) A.Weber & D.J.Middleton, Taxon 60: 778 (2011). 
Chirita micromusa B.L.Burtt, J. Roy. Hort. Soc. 85: 28 (1960); Barnett, Fl. Siam. 3: 226 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 194 (1974); Burtt, Thai Forest Bull., Bot. 29: 88 (2001). 

– TYPE: Thailand, Nakhon Nayok, cult. in Montreal Botanic Garden from seeds collected by Raymond & Smitinand, Raymond, M. ref. 106-59 (holotype E [E00155279]).

18. Microchirita mollissima (Ridl.) A.Weber & D.J.Middleton, Taxon 60: 778 (2011).
 – Chirita mollissima Ridl., J. Linn. Soc., Bot. 32: 517 (1896); Barnett, Fl. Siam. 3: 226 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 188 (1974); Burtt, Thai Forest Bull., Bot. 29: 88 (2001). 

– TYPE: Siam [Thailand], Pungah [Phangnga], July 1893, Curtis 2944 (lectotype SING [SING0117733] ....

18a. Microchirita mollissima var. mollissima 
18b. Microchirita mollissima var. glabra C.Puglisi, var. nov. 
18c. Microchirita mollissima var. glandulophylla C.Puglisi, var. nov. 


19. Microchirita oculata (Craib) A.Weber & D.J.Middleton, Taxon 60: 778 (2011).
 – Chirita oculata Craib, Bull. Misc. Inform. Kew. 174 (1930); Barnett, Fl. Siam. 3: 226 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 194 (1974); Burtt, Thai Forest Bull., Bot. 29: 88 (2001). 

– TYPE: “described from a plant which flowered in Aberdeen in Jul 1928. It was raised from seed of Kerr 9750 which was collected on Kao Sakan, 6 November 1928” (lectotype ABD, designated by Wood (1974: 194); isolectotype E [E00155281]). (Fig. 12D–F)


20. Microchirita personata C.Puglisi, Kew Bull. 71(1)-2: 1 (2016). 
– TYPE: Thailand, Uthai Thani, Lan Sak, Huppatat Non Hunting Area, 122 m, 14 October 2014, Middleton, D.J., Hemrat, C., Karaket, P., Puglisi, C. & Suddee, S. 5688 (holotype BKF; isotypes AAU, E [E00663026], K, SING). (Fig. 13A–C)

21. Microchirita purpurea D.J.Middleton & Triboun, Thai Forest Bull., Bot. 41: 14 (2013). 
– TYPE: Thailand, Chanthaburi, Kaeng Hang Maeo, Khao Chamao National Park, ..., 30 m alt., 27 August 2012, Middleton, D.J., Karaket, P., Suddee, S. & Triboun, P. 5681 (holotype BKF; isotypes A [00435722], BK, E [E00626983], K, P [P00966760], QBG, SING [SING0229833]). 



22. Microchirita rupestris (Ridl.) A.Weber & Rafidah, Taxon 60: 779 (2011); Rafidah, Gard. Bull. Singapore 69: 18 (2017). – Chirita rupestris Ridl., J. Straits Branch Roy. Asiat. Soc. 44: 59 (1905); Barnett, Fl. Siam. 3: 227 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 201 (1974); Burtt, Thai Forest Bull., Bot. 29: 89 (2001). – TYPE: Malaysia, Kedah, Langkawi, on damp rocks, sea level, November 1889, Curtis 2120 (lectotype SING [SING0042989], designated by Puglisi in Rafidah (2017: 18); isolectoype SING [SING0042990]). (Fig. 14)


23. Microchirita suddeei D.J.Middleton & Triboun, Thai Forest Bull., Bot. 41: 18 (2013). – TYPE: Thailand, Phrae, Rong Kwang District, Tham Pha Nang Khoi, 210 m alt., 17 August 2012, Middleton, D.J., Karaket, P., Suddee, S. & Triboun, P. 5618 (holotype BKF; isotypes E [E00629451], P [P00966761], QBG, SING [SING0229832]). (Fig. 15A–C)


  

หยาดตาดโพธิ์  Microchirita tadphoensis C. Puglisi (Gesneriaceae) จากอุทยานแห่งชาติภูลังกา อำเภอบ้านแพง จังหวัดหนองนครพนม ไม้ล้มลุกขึ้นตามก้อนหินทรายที่ชื้น ลำต้นอวบน้ำ ดอกสีเหลือง กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก 


24. Microchirita tadphoensis C.Puglisi, sp. nov.  
Most similar to Microchirita hamosa (R.Br.) Yin Z.Wang in the delicate habit and to M. bimaculata (D.Wood) A.Weber & D.J.Middleton in the shape of the corolla. Differs in having a shortly campanulate pale yellow corolla (white in Microchirita hamosa) with a ventral darker yellow marking but no lateral spots (spots always present in M. bimaculata).
 – TYPE: Thailand, Nakhon Phanom, Ban Phaeng, Phu Langka National Park, Tad Pho Waterfall, 224 m, 23 October 2015, Suddee, S., Keiwbang, W. & Hemrat, C. 4980 (holotype BKF; isotype SING). (Fig. 15D–F)


Fig. 16. Microchirita tetsanae C.Puglisi. A. Habit. B. Front view of the flower. All from Tetsana, N. et al. 855.
Microchirita thailandica C.Puglisi. C. Habit. D. Front view of the flower. E. Lateral view of the flower. All from Tetsana, N. et al. 904. (Photos: N. Tetsana)

    

ม่วงเทศนา Microchirita tetsanae C. Puglisi (Gesneriaceae) 
พรรณไม้ถิ่นเดียวของไทย (endemic) จากเขาหินปูน อำเภอเมือง จังหวัดเพชรบูรณ์ ไม้ล้มลุกขึ้นตามหน้าผาหินปูน ลำต้นอวบน้ำ ดอกสีม่วง กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก แฉกบน 2 กลีบ โคนไม่ซ้อนทับแฉกล่าง โคนกลีบปากมีแต้มสีเหลือง 

25. Microchirita tetsanae C.Puglisi, sp. nov.
 Species characterised by the presence of a dimorphic indumentum on the anthers and sparse hairs on the filaments, and by a little projection at the anther insertion. It is most similar to Microchirita thailandica C.Puglisi, but differs in the upper lobes not being imbricate with the lower, and in the filament projection. – TYPE: Thailand, Phetchabun, Mueang Phetchabun, Wat Tham Nam Bang, 130 m, 13 September 2014, Tetsana, N. et al. 855 (holotype BKF; isotype SING). (Fig. 16A–B)


26. Microchirita thailandica C.Puglisi, sp. nov.
 Species most similar to Microchirita tetsanae C.Puglisi in the colour pattern of the corolla, differing in having a narrower tube which widens abruptly (gradually widening in M. tetsanae), all corolla lobes imbricate (vs. lateral lobes not imbricate with the upper in M. tetsanae), a shorter ventral tube, and in not having a projection at the anther insertion. – TYPE: Thailand, Chaiyaphum, Phak Dee Chumphon, Wat Thum Wua Daeng, 460 m, 8 November 2014, Tetsana, N. et al. 904 (holotype BKF; isotype SING). (Fig. 16C–E)

27. Microchirita tubulosa (Craib) A.Weber & D.J.Middleton, Taxon 60: 79 (2011). 

Chirita tubulosa Craib, Bull. Misc. Inform. Kew 173 (1922); Barnett, Fl. Siam. 3: 227 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 190 (1974); Burtt, Thai Forest Bull., Bot. 29: 89 (2001). 

– TYPE: “described from plants grown from seed collected by Kerr. Flowered in October 1921” (lectotype ABD, first step designated by Wood (1974: 190), second sted designated here [the specimen which matches the image deposited in E, barcode E00155288]; isolectotypes ABD, E). (Fig. 17)


28. Microchirita viola (Ridl.) A.Weber & Rafidah, Taxon 60: 779 (2011); Rafidah, Gard. Bull. Singapore 69: 26 (2017).
– Chirita viola Ridl., J. Linn. Soc., Bot. 32: 516 (1896); Barnett, Fl. Siam. 3: 228 (1962); Wood, Notes Roy. Bot. Gard. Edinburgh 33: 190 (1974); Burtt, Thai Forest Bull., Bot. 29: 89 (2001). – Didymocarpus viola (Ridl.) Williams, Bull. Herb. Boiss. Ser. 2, 5: 434 (1905).

– TYPE: Malaysia, Peninsular Malaysia, Kedah, Langkawi, September 1890, Curtis 2570 (lectotype SING [SING0042993], designated by Rafidah (2017: 26); isolectotypes SING [SING0042994, SING0042995]). (Fig. 18A–B)



29. Microchirita woodii D.J.Middleton & Triboun, Thai Forest Bull., Bot. 41: 15 (2013).
– TYPE: Thailand, Nan, Muang Nan, Tham Pha Tup Forest Park, trail to Phra Cave, 300 m alt., 16 August 2012, Middleton, D.J., Karaket, P., Suddee, S. & Triboun, P. 5612 (holotype BKF; isotypes BK, E [00629450], P [P00966763]).


   

C. Puglisi and D.J. Middleton. 2017.  A Revision of Microchirita (Gesneriaceae) in Thailand. Gardens' Bulletin Singapore. 69(2); 211 - 284. 


  

หยาดผาผึ้ง Microchirita limbata C. Puglisi (Gesneriaceae) 
จากเขตรักษาพันธุ์สัตว์ป่าผาผึ้ง อำเภอคอนสาร จังหวัดชัยภูมิ ไม้ล้มลุกขึ้นตามก้อนหินปูน ลำต้นอวบน้ำ ดอกสีม่วง กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก 
เป็นพรรณไม้ชนิดใหม่ของโลกและเป็นพรรณไม้ถิ่นเดียวของไทย (endemic) ตีพิมพ์ในวารสาร Gardens' Bulletin Singapore 69(2): 211–284. ตัวอย่างต้นแบบ Suddee, Keiwbang & Hemrat 4968 เก็บจากเขาหินปูน ถ้ำฮวงโป เขตรักษาพันธุ์สัตว์ป่าผาผึ้ง 

    

ม่วงเทศนา Microchirita tetsanae C. Puglisi (Gesneriaceae) 
จากเขาหินปูน อำเภอเมือง จังหวัดเพชรบูรณ์ ไม้ล้มลุกขึ้นตามหน้าผาหินปูน ลำต้นอวบน้ำ ดอกสีม่วง กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก แฉกบน 2 กลีบ โคนไม่ซ้อนทับแฉกล่าง โคนกลีบปากมีแต้มสีเหลือง
พรรณไม้ชนิดใหม่ของโลกและเป็นพรรณไม้ถิ่นเดียวของไทย (endemic) ตีพิมพ์ในวารสาร Gardens' Bulletin Singapore 69(2): 211–284. 2017 คำระบุชนิด ‘tetsanae’ มาจากนามสกุล ดร.นัยนา เทศนา ผู้เก็บตัวอย่าง Tetsana et al. 855 ตัวอย่างต้นแบบแรก (holotype) เก็บรักษาไว้ที่หอพรรณไม้ (BKF)

  


หยาดสวนสวรรค์ Microchirita hypocrateriformis C. Puglisi จากเขาหินปูน อำเภอเอราวัณ จังหวัดหนองบัวลำพู ไม้ล้มลุกขึ้นตามเขาหินปูน ลำต้นอวบน้ำ ดอกสีเหลืองอ่อนหรือสีม่วง แต่ที่พบมากจะเป็นสีเหลืองอ่อน กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก 
พรรณไม้ชนิดใหม่ของโลกและเป็นพรรณไม้ถิ่นเดียวของไทย (endemic) ตีพิมพ์ในวารสาร Gardens' Bulletin Singapore 69(2): 211–284. พบขึ้นตามเขาหินปูนในภาคตะวันออกเฉียงเหนือของไทย เช่นบริเวณสวนสวรรค์ สวนหินผางาม อำเภอหนองหิน จังหวัดเลย ตัวอย่างต้นแบบ Suddee, Keiwbang & Hemrat 4967 เก็บจากวัดถ้ำฮวงโป อำเภอคอนสาร จังหวัดชัยภูมิ

  

  

หยาดตาดโพธิ์ Microchirita tadphoensis C. Puglisi (Gesneriaceae) จากอุทยานแห่งชาติภูลังกา อำเภอบ้านแพง จังหวัดหนองนครพนม ไม้ล้มลุกขึ้นตามก้อนหินทรายที่ชื้น ลำต้นอวบน้ำ ดอกสีเหลือง กลีบดอกเชื่อมกันเป็นหลอด ปลายแยกเป็น 5 แฉก
พรรณไม้ชนิดใหม่ของโลกและเป็นพรรณไม้ถิ่นเดียวของไทย (endemic) ตีพิมพ์ในวารสาร Gardens' Bulletin Singapore 69(2): 211–284. ตัวอย่างต้นแบบ Suddee, Keiwbang & Hemrat 4980 เก็บจากป่าดิบแล้ง เส้นทางน้ำตกตาดโพธิ์ อุทยานแห่งชาติภูลังกา

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Mammalogy • 2018] Cassistrellus gen. nov. • A New Genus and Species of Vespertilionid Bat from the Indomalayan Region: Cassistrellus dimissus & C. yokdonensis ---ScRaBBlE


Cassistrellus dimissus (Thomas, 1916)

Ruedi, Eger, Lim & Csorba, 2018

Abstract
Bats belonging to the subfamily Vespertilioninae are diverse and cosmopolitan, but their systematic arrangement remains a challenge. Previous molecular surveys suggested new and unexpected relationships of some members compared to more traditional, morphology-based classifications, and revealed the existence of taxonomically undefined lineages. We describe here a new genus and species corresponding to an enigmatic lineage that was previously identified within the genus Eptesicus in the Indomalayan Region. Phylogenetic reconstructions based on mitochondrial and nuclear genes relate the new taxon to Tylonycteris and Philetor, and show that specimens associated with this new genus represent 2 genetically distinct species. Although little is known about their ecology, locations of capture and wing morphology suggest that members of this new genus are tree-dwelling, open-space aerial insect predators. The new species has only been documented from Yok Don National Park in Vietnam, so its conservation status is uncertain until more surveying methods target the bat fauna of the dipterocarp forest in Southeast Asia.

Key words: eptesicoid bats, phylogeny, systematics, taxonomy, Vespertilionidae 

Fig. 1. Maximum likelihood (ML) phylogeny of selected Vespertilioninae bats based on a combined alignment of 2 mitochondrial and 1 nuclear gene (2,161 bp in total). Numbers near nodes indicate branch support (left, percent ML bootstrap, right, posterior probability), while solid circles represent nodes recovered in >98% of both resampling methods. The gray box highlights the strongly supported clade comprising species of Cassistrellus gen. nov., Tylonycteris, and Philetor

Fig. 2. Portrait of a live adult male Cassistrellus dimissus captured near the Royal Chitwan National Park in Nepal in March 1990. This specimen was collected as part of the series of vouchers described by Myers et al. (2000b) and conserved at the UMMZ
(photo: Phil Myers).  DOI: 10.1093/jmammal/gyx156

 Fig. 3. Dorsal, ventral, and lateral views of the skull and mandible of a male Cassistrellus dimissus from Laos (MHNG 1926.053). Notice the deep basisphenoid pits between tympanic bullae, long, cuspidate upper canines, strong lambdoidal and occipital crests, and prominent preorbital processes that are typical morphological features of Cassistrellus gen. nov.

Cassistrellus gen. nov.

Type species: Eptesicus dimissus Thomas, 1916

Description: Species of Cassistrellus are medium-sized vespertilionids (FA 39–47 mm; body mass 12–17 g) characterized externally by short, chestnut-brown pelage that is paler on the ventral parts, by narrow wings with short and pointed tips, and an especially broad muzzle (Fig. 2). Wing membranes are attached to the middle or distal parts of the metatarsus. The tail is mostly included in the uropatagium and extends by 2–3 mm beyond its posterior margin. The calcar extends less than halfway to the tail and may have a small lobe near the ankle. The skull is robust and angular in profile making an almost straight line between the rostrum and the occipital region. It is characterized by well-developed sagittal and lambdoid crests, which meet near the top of the skull to form an occipital helmet. On its ventral surface, the skull has a pair of deep and well-delimited basisphenoid pits located between the cochleae (Fig. 3). Laterally, the lachrymal region has prominent preorbital processes, but the supraoccipital ridges are weak and barely visible. The dental formula is  2113/3123 = 32  teeth, with the upper canines possessing a distinct secondary cusp along the rear edge (Fig. 3).

Etymology: The name Cassistrellus derives from the Latin noun “cassis”, which means wearer of a helmet, in reference to the shape of the hind parts of the skull. The suggested English vernacular name is helmeted bat.

Geographic distribution: The type specimen of C. dimissus was collected by H. C. Robinson and E. Seimund in Kao Nawng, Bandon (currently within Tai Rom Yen National Park in Surat Thani province of Thailand) at 1,400 feet (= 436 m a.s.l.), in June 1913 (Robinson and Kloss 1915). However, the altitude associated with this specimen was reported by Thomas (1916) as 3,500 feet (= 1,067 m a.s.l.), which would correspond to near the summit of the Khao Nong mountain, where the collectors did not capture bats. As all known localities of Cassistrellus are located in the lowlands at elevation between 190 and 674 m a.s.l., these bats should be indeed regarded as lowland dwellers. The vast area covered by the few scattered records of Cassistrellus suggests that it should be widely distributed from the Isthmus of Kra into mainland Southeast Asia and the foothills of the Himalaya, i.e., across most of the Indo-Burma biodiversity hotspot (Myers et al. 2000a). All capture sites were situated in hilly terrain with mixed deciduous or dipterocarp forests traversed by large rivers.

Fig. 4. Lateral view of skull of ACassistrellus yokdonensis sp. nov. (holotype, ROM 107751) and BC. dimissus (holotype, BM(NH) 16.4.21.1.). The scale bar at the bottom represents 5 mm. 


Fig. 4. Lateral view of skull of CC. dimissus from Laos (MHNG 1926.053), and DC. dimissus from Nepal (UMMZ 172218). The scale bar at the bottom represents 5 mm. 


Fig. 4. Lateral view of skull of A) Cassistrellus yokdonensis sp. nov. (holotype, ROM 107751), B) C. dimissus (holotype, BM(NH) 16.4.21.1.), C) C. dimissus from Laos (MHNG 1926.053), and D) C. dimissus from Nepal (UMMZ 172218).
The scale bar at the bottom represents 5 mm. 

Cassistrellus yokdonensis sp. nov.
Yok Don helmeted bat
Eptesicus sp. A JLE-2010: Francis et al., 2010

Holotype: Male ROM 107751 (field number 42734) collected on 6 June 1997 by B. K. Lim and M. Theberge. Preserved as a skin, skull, and partial skeleton. Epiphyses almost completely fused, indicating that this bat was a subadult.

Type locality: Vietnam: Dak Lak province; Yok Don National Park, Dak Ken River (tributary of the Serepok River), base of Yok Mt. ..., at 194 m a.s.l. in dry, open dipterocarp forest.

Diagnosis: Cassistrellus yokdonensis sp. nov. is a medium-sized vespertilionid bat (body mass about 15 g) characterized by pointed, narrow wings similar in shape to those of Nyctalus species, but not as narrow. The fur is sparse with short hairs, clove brown (Ridgway 1912) dorsally, lighter beige ventrally, and cream colored at the throat. There is no glossy tinge to the fur. The color of the wings and other skin parts is blackish brown. The wing membranes attach to the distal end of the metatarsus. The calcar extends less than halfway to the tail and has no visible lobe.

Although the dental formula is identical to that of Eptesicus species, the skull possesses a pair of deep and well-defined basisphenoid pits and prominent preorbital processes that are otherwise absent from the latter genus. C. yokdonensis sp. nov. is morphologically similar to C. dimissus, but—in spite of the fact that the known individuals are not fully grown adults—is substantially larger, both externally (e.g., FA 47 mm versus 39–42 mm; Table 1), and cranially (e.g., maxillary toothrow length over 6.5 mm versus less than 6.4 mm). C. yokdonensis sp. nov. has also a much stronger dentition in general, and subequal small lower premolars, longer more curved upper canines, and procumbent upper incisors compared to its congener. Genetically, C. yokdonensis sp. nov. has unique mitochondrial (Cytb, Co1) and nuclear (Rag2) sequences compared to C. dimissus from Laos.

Etymology: We propose the name C. yokdonensis after the national park where it has been found, in recognition of the importance of protected areas in conserving species and their habitats.

Geographic distribution: Currently known only from Yok Don National Park, Dak Lak Province of Vietnam. The 2 specimens were caught shortly after 2100 h as they flew into large (30 m long by 10 m high) canopy nets deployed in a dry, open dipterocarp forest of lowland regions. Other species of mammals caught in the same nets included Taphozous, Pipistrellus, Murina, Cynopterus, Megaerops, and Rhinolophus bats and several Hylopetes flying squirrels. In addition, a paratype specimen of a new species of parachute gecko (Ptychozoon trinitaterra) was caught in this net (Brown 1999).


Manuel Ruedi, Judith L. Eger, Burton K. Lim and Gábor Csorba. 2017. A New Genus and Species of Vespertilionid Bat from the Indomalayan Region. Journal of Mammalogy. gyx156. DOI: 10.1093/jmammal/gyx156

   

    

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

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