The new species Dregea taynguyenensis from Vietnam is described, illustrated and compared with the other Dregea species occurring in Vietnam and neighbouring countries D. cuneifolia, D. sinensis, D. volubilis and D. yunnanensis. Dregea taynguyenensis differs from these species by the leaf blade secondary veins number, calyx lobes shape, and corolla lobes size.
Dregea taynguyenensis is most similar to D. cuneifolia due to the elliptic-oblong leaf blades, cuneate at base and acuminate at apex. The former is distinguished from the latter by having 7–10 secondary veins each side of midrib (vs. 5–6 in D. cuneifolia), calyx lobes lanceolate to linear (vs. ovate) and corolla lobes 10.2–12.6 × 5.3–6.4 mm (vs. c. 4 × 4 mm). ....
Etymology:— The specific epithet refers to the type locality, Tay Nguyen area in Vietnam..
Distribution and ecology:— Dregea taynguyenensis is only found in Vietnam, Gia Lai province, Tay Nguyen area where it grows in sunny places in secondary forest, in association with Bidens pilosa Linnaeus (1753: 832), Jasminum lanceolarium Roxburg (1820: 97), Maclurodendron oligophlebium (Merrill 1923: 246) T.G. Hartley (1982: 13), Mallotus paniculatus (Lamarck in Lamarck & Poiret 1786: 207) Müller Argoviensis (1865: 34), Melicope pteleifolia (Champion ex Bentham in Bentham 1851: 330) T.G. Hartley (1993: 521).
FIGURE 1. Dregea taynguyenensis. Flowering branch in the habitat of the type locality. Photograph by T.B. Tran (9 June 2011). FIGURE 2. Dregea taynguyenensis.A, Leaf blade, abaxial side (left), adaxial side (right); B, Inflorescence with flower buds; C, Flowering branch; D, Inflorescence with greenish flowers; E, Inflorescence with flowers yellowish flushed purple. Based on T.B. Tran et al. Bach 0906201101 (HN), photographs by T.B. Tran (9 June 2011).
The Bach Tran, Hai Do Van, Ha Bui Thu, Sangho Choi, Sangmi Eum and Michele Rodda. 2018. Dregea taynguyenensis (Apocynaceae, Asclepiadoideae), A New Species from Vietnam. Phytotaxa. 333(2); 267–273. DOI: 10.11646/phytotaxa.333.2.11
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
We describe a new species of Rhacophorus based on two adult specimens collected from Hoa Binh Province, northern Vietnam. Rhacophorus hoabinhensis sp. nov. is distinguishable from its congeners on the basis of a combination of the following morphological characters: size small (SVL 31.1–32.5 mm in males); head slightly longer than wide; vomerine teeth absent; snout short (SNL/SVL 0.16); dorsal skin smooth; forearm and tarsus with dermal fringes; dermal appendage at vent present; webbing formula on fingers I12/3-12/3II1-2III1-1IV and on toes I3/4-1II1/2- 1III1/2-1IV1-1/2V; dorsal surface grey yellow with brown spots; lower jaw region dark grey, throat, chest and belly cream; anterior and posterior thighs, as well as ventral surface of tibia orange. The interspecific uncorrected genetic distances (16S rRNA gene) between the new species from Hoa Binh and other analyzed congeners varied from 9.8% to 17.4%. In the phylogenetic analyses, the new species revealed to be a representative of Rhacophorus and was nested within the R. hoanglienensis-orlovi species group.
Keywords:Rhacophorus hoabinhensis sp. nov., karst forest, molecular phylogeny, taxonomy, Hoa Binh Province
Figure 2 Dorsal and ventral views of the adult male holotype (IEBR A.2016.18) ofRhacophorus hoabinhensis sp. nov. from Hoa Binh Province, northern Vietnam.
Figure 4 Rhacophorushoabinhensis sp. nov. in its biotope in Hang Kia–Pa Co Nature Reserve, Hoa Binh Province, Vietnam.
Rhacophorus hoabinhensis sp. nov.
Etymology: The specific epithet “hoabinhensis” refers to the type locality of the new species, Hoa Binh Province. For the common names we suggest Hoa Binh Treefrog(English), and Ếch cây hòa bình (Vietnamese).
Distribution:R. hoabinhensis is currently is known only from the type locality in Hang Kia–Pa Co Nature Reserve, Hoa Binh Province, Vietnam.
Natural history:Rhacophorus hoabinhensis appears to be closely associated with karstic environment. Specimens were found at night between 19:00 and 23:30h, near a small pond. The surrounding habitat was secondary karst forest, consisting of medium and small hardwoods mixed with shrubs and vines. Specimens were found on leaves and branches of trees, about 1.2–1.5 m above the ground. The advertisement call of the species was not heard. The air temperatures at the times of collection ranged from 14.8 to 18.9o C and relative humidity from 84% to 93%. Other rhacophorid species recorded in Hang Kia–Pa Co Nature Reserve were Raorchestes parvulus Boulenger, R. feaeBoulenger, R. kio Ohler and Delorme, R. orloviZiegler and Köhler, Theloderma albopunctatumLiu and Hu, T. gordoniTaylor, and T. lateriticum Bain, Nguyen and Doan. Females and the tadpole of Rhacophorus hoabinhensis have not been recorded so far.
Tao Thien Nguyen, Cuong The Pham, Truong Quang Nguyen, Hoa Thi Ninh and Thomas Ziegler. 2017. A New Species of Rhacophorus (Amphibia: Anura: Rhacophoridae) from Vietnam. Asian Herpetological Research.8(4); 221–234. DOI: 10.16373/j.cnki.ahr.170046
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
Abstract Drepanosticta emtrai sp. nov. is described from Vietnam (holotype male Ha Tinh Province, 9 vi 2015, to be deposited in RMNH). The new species is allied to D. carmichaeli(Laidlaw, 1915) and a number of other species of Drepanosticta including D. vietnamicaAsahina, 1997. New illustrations of the paratype of D. vietnamica are provided and the species is discussed. The Drepanostictacarmichaeli-group, to which the above mentioned species belong, is defined and discussed.
Keywords: Odonata, Platystictidae, Vietnam, Drepanosticta emtrai, Drepanosticta carmichaeli-group, Drepanosticta vietnamica, new species
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
Asian Mountain Toads (Ophryophryne) are a poorly known genus of mostly small-sized anurans from southeastern China and Indochina. To shed light on the systematics within this group, the most comprehensive mitochondrial DNA phylogeny for the genus to date is presented, and the taxonomy and biogeography of this group is discussed. Complimented with extensive morphological data (including associated statistical analyses), molecular data indicates that the Langbian Plateau, in the southern Annamite Mountains, Vietnam, is one of the diversity centres of this genus where three often sympatric species of Ophryophryne are found, O. gerti, O. synoria and an undescribed species. To help resolve outstanding taxonomic confusion evident in literature (reviewed herein), an expanded redescription of O. gerti is provided based on the examination of type material, and the distributions of both O. gerti and O. synoria are considerably revised based on new locality records. We provide the first descriptions of male mating calls for all three species, permitting a detailed bioacoustics comparison of the species. We describe the new species from highlands of the northern and eastern Langbian Plateau, and distinguish it from its congeners by a combination of morphological, molecular and acoustic characters. The new species represents one of the smallest known members of the genus Ophryophryne. At present, the new species is known from montane evergreen forest between 700–2200 m a.s.l. We suggest the species should be considered Data Deficient following IUCN’s Red List categories.
Keywords: 12S rRNA, 16S rRNA, advertisement call, amphibian, biodiversity, Da Lat Plateau, frog, Indochina, southeast Asia, taxonomy, Truong Son
Figure 13. Comparison of the head coloration in life of three Ophryophryne species from the Langbian Plateau: A Ophryophryne synoria, Bu Gia Map N.P., Binh Phuoc Prov., Vietnam B Ophryophryne gerti, Chu Yang Sin N.P., Dak Lak Prov., Vietnam C Ophryophryne elfina sp. n., Hon Giao Mt., Bidoup–Nui Ba N.P., Lam Dong Prov. Photos by N.A. Poyarkov and N.L. Orlov.
Figure 6. Ophryophryne species of the Langbian Plateau (Vietnam): AOphryophryne gerti, female, Chu Yang Sin N.P., Dak Lak Prov., 1000 m a.s.l. B O. gerti, male, Chu Yang Sin N.P., Dak Lak Prov., 1000 m a.s.l. CO. gerti, male, Bidoup–Nui Ba N.P., Lam Dong Prov., 1550 m a.s.l. DO. synoria, female, Bu Gia Map N.P., Binh Phuoc Prov., 400 m a.s.l. EO. synoria, male, Chu Yang Sin Mt., Chu Yang Sin N.P., Dak Lak Prov., 1000 m a.s.l. FO. synoria, male, Bidoup–Nui Ba N.P., Lam Dong Prov., 1550 m a.s.l. G Ophryophryne elfina sp. n., male, Chu Yang Sin Mt., Chu Yang Sin N.P., Dak Lak Prov., 2000 m a.s.l. HOphryophryne elfina sp. n., male, Hon Giao Mt., Bidoup–Nui Ba N.P., Lam Dong and Khanh Hoa provincial border, 2000 m a.s.l.
Photos by N.A. Poyarkov and N.L. Orlov.
• Taxonomic remarks on Ophryophryne gerti Ohler, 2003
Redescription of the holotype: Mature male (SVL 35.7 mm), habitus slender (Fig. 7A, B). Specimen in good state of preservation; two incisions are present on trunk, one longitudinally orientated on mid-abdomen, another longitudinally oriented on upper flank on right side; liver and testes observable through incisions, testes enlarged; jaw is dislocated on right allowing visual access to buccal cavity. .....
• New records and range extension for Ophryophryne synoriaStuart, Sok & Neang, 2006
Distribution and remarks: Stuart et al. (2006) described a large-sized Ophryophryne from O’Rang (also spelled as “O’Reang”) District in eastern Cambodia, close to the Vietnamese border, as O. synoria (Loc. 1, Fig. 1). Subsequently, during field surveys in 2009–2011, the species was reported in southern Vietnam from Bu Gia Map N.P., Binh Phuoc Prov. (Loc. 2, Fig. 1) and Cat Tien N.P. in Dong Nai Prov. (Loc. 3, Fig. 1) based on morphological evidence (Vassilieva et al. 2016). Herein, we confirm the identity of these specimens based on morphological and molecular genetic evidence, and further expand its distribution in southern Vietnam to include medium and low elevation localities in the central and western parts of the Langbian Plateau (Dak Lak, Lam Dong, Dong Nai and Binh Phuoc provinces between 200 and 1500 m a.s.l.; its presence in Dak Nong Prov. is anticipated). We also identify two mtDNA lineages within O. synoria with a moderate level of sequence divergence (p = 2.6%: Table 2, Fig. 2): Subclade B inhabits mountain areas in Lam Dong and Dak Lak provinces and was also recorded for the lowland habitat in Dong Nai Prov. (Locs. 3–4, 6 and 11, Fig. 1) whereas Subclade C is only found in Mondolkiri Prov. of Cambodia and adjacent Binh Phuoc Prov. of Vietnam (Fig. 1, Locs 1–2) and corresponds to O. synoria s. stricto.
Figure 14. Ophryophryne elfina sp. n.in situ: A Two syntopically collected males of Ophryophryne gerti (left) and Ophryophryne elfina sp. n. (right) in Chu Yang Sin N.P., Dak Lak Prov., Vietnam, 1750 m a.s.l., photo by N.L. Orlov B calling adult male of Ophryophryne elfina sp. n. in Nui Chua Mt., Nui Chua N.P., Ninh Thuan Prov., Vietnam, 780 m a.s.l., photo by S.N. Nguyen C adult male of Ophryophryne elfina sp. n. in calling position in Hon Ba N.R., Khanh Hoa Prov., Vietnam, 1510 m a.s.l., photo by L.T. Nguyen.
• Description of a new species of Ophryophryne
Based upon several lines of evidence, including the analyses of diagnostic morphological characters, acoustic analyses of advertisement calls and phylogenetic analyses of mtDNA sequences for the 12S rRNA–16S rRNA genes, the new species of Ophryophryne from mid to high elevations of the western Langbian Plateau represents a highly divergent mtDNA lineage, clearly distinct from all other Ophryophryne species. These results support our hypothesis that this recently discovered lineage of Ophryophryne represents an undescribed species, described below:
Ophryophryne elfina sp. n.
Etymology: The specific epithet is an adjective (in agreement with the genus name in feminine gender), derived from “elf”, the English spelling of “alfus” in Latin, referring to usually forest-dwelling supernatural mythological creatures in Germanic mythology and folklore; the name is given in reference both to the funny appearance and small size of the new species, as well as to the their endangered habitat, restricted to wet evergreen montane forests at high elevations of the Langbian Plateau; such forests are often called “elfin forests”.
Recommended vernacular name: The recommended common name in English is “Elfin Mountain Toad”. The recommended common name in Vietnamese is “Cóc Núi Tiểu Yêu Tinh”.
Figure 9. Holotype of Ophryophryne elfina sp. n. in life (ZMMU A-5669, male, field number NAP-02658), dorsolateral view. Photos by N.A. Poyarkov.
Figure 10. Paratypes ofOphryophryne elfina sp. n. in life. A–D Bidoup Mt., Bidoup–Nui Ba N.P., Lam Dong Prov., 2000 m a.s.l.: A ZMMU A-4788 (field number NAP-01449), male, dorsolateral view B ZMMU A-4788 (field number NAP-01455), female, dorsolateral view C ZMMU A-4788 (field number NAP-01449), male, ventral view D ZMMU A-4788 (field number NAP-01455), female, ventral view E–F Chu Yang Sin Mt., Chu Yang Sin N.P., Dak Lak Prov., 1800 m a.s.l.: E ZMMU A-5691 (field number ABV-00580), metamorph, dorsolateral view F ZMMU A-5691 (field number ABV-00581), metamorph, dorsolateral view. Photos by N.A. Poyarkov.
Distribution: Ophryophryne elfina sp. n. is found to be endemic to five provinces in (Lam Dong, Dak Lak, Khanh Hoa, Ninh Thuan and Phu Yen) in the northern and eastern part of the Langbian Plateau and its foothills in southern Vietnam (localities 6–12, Fig. 1). The new species is restricted to wet evergreen montane tropical and elfin forests, receiving high precipitation from the sea. Such wet forests are found only on high elevations in the central parts of the Langbian Plateau (e.g. 1900–2100 m a.s.l. on Bidoup Mt., Lam Dong Prov., Fig. 1, Loc. 6) or peripheral mountains remote from the sea (e.g. 1900–2300 m a.s.l. on Chu Pan Fan and Chu Yang Sin Mts., Dak Lak Prov., Fig. 1, Locs 10 and 11), but on the eastern foothills of the plateau which receive more precipitation, the new species is found at lower elevation (from 950 to 1510 m a.s.l. on Hon Ba Mt., Khanh Hoa Prov., Fig. 1, Loc. 8; 780 m a.s.l. on Nui Chua Mt., Ninh Thuan Prov., Fig. 1, Loc. 9; and 700 m in Phu Yen Prov., Fig. 1, Loc. 12).
.....
Nikolay A. Poyarkov Jr., Tang Van Duong, Nikolai L. Orlov, Svetlana S. Gogoleva, Anna B. Vassilieva, Luan Thanh Nguyen, Vu Dang Hoang Nguyen, Sang Ngoc Nguyen, Jing Che and Stephen Mahony. 2017. Molecular, Morphological and Acoustic Assessment of the Genus Ophryophryne (Anura, Megophryidae) from Langbian Plateau, southern Vietnam, with Description of A New Species. ZooKeys. 672: 49-120. DOI: 10.3897/zookeys.672.10624
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
Bats belonging to the subfamily Vespertilioninae are diverse and cosmopolitan, but their systematic arrangement remains a challenge. Previous molecular surveys suggested new and unexpected relationships of some members compared to more traditional, morphology-based classifications, and revealed the existence of taxonomically undefined lineages. We describe here a new genus and species corresponding to an enigmatic lineage that was previously identified within the genus Eptesicus in the Indomalayan Region. Phylogenetic reconstructions based on mitochondrial and nuclear genes relate the new taxon to Tylonycteris and Philetor, and show that specimens associated with this new genus represent 2 genetically distinct species. Although little is known about their ecology, locations of capture and wing morphology suggest that members of this new genus are tree-dwelling, open-space aerial insect predators. The new species has only been documented from Yok Don National Park in Vietnam, so its conservation status is uncertain until more surveying methods target the bat fauna of the dipterocarp forest in Southeast Asia.
Fig. 1. Maximum likelihood (ML) phylogeny of selected Vespertilioninae bats based on a combined alignment of 2 mitochondrial and 1 nuclear gene (2,161 bp in total). Numbers near nodes indicate branch support (left, percent ML bootstrap, right, posterior probability), while solid circles represent nodes recovered in >98% of both resampling methods. The gray box highlights the strongly supported clade comprising species of Cassistrellus gen. nov., Tylonycteris, and Philetor.
Fig. 2. Portrait of a live adult male Cassistrellus dimissus captured near the Royal Chitwan National Park in Nepal in March 1990. This specimen was collected as part of the series of vouchers described by Myers et al. (2000b) and conserved at the UMMZ
Fig. 3. Dorsal, ventral, and lateral views of the skull and mandible of a male Cassistrellus dimissus from Laos (MHNG 1926.053). Notice the deep basisphenoid pits between tympanic bullae, long, cuspidate upper canines, strong lambdoidal and occipital crests, and prominent preorbital processes that are typical morphological features of Cassistrellus gen. nov.
Description: Species of Cassistrellus are medium-sized vespertilionids (FA 39–47 mm; body mass 12–17 g) characterized externally by short, chestnut-brown pelage that is paler on the ventral parts, by narrow wings with short and pointed tips, and an especially broad muzzle (Fig. 2). Wing membranes are attached to the middle or distal parts of the metatarsus. The tail is mostly included in the uropatagium and extends by 2–3 mm beyond its posterior margin. The calcar extends less than halfway to the tail and may have a small lobe near the ankle. The skull is robust and angular in profile making an almost straight line between the rostrum and the occipital region. It is characterized by well-developed sagittal and lambdoid crests, which meet near the top of the skull to form an occipital helmet. On its ventral surface, the skull has a pair of deep and well-delimited basisphenoid pits located between the cochleae (Fig. 3). Laterally, the lachrymal region has prominent preorbital processes, but the supraoccipital ridges are weak and barely visible. The dental formula is 2113/3123 = 32 teeth, with the upper canines possessing a distinct secondary cusp along the rear edge (Fig. 3).
Etymology: The name Cassistrellus derives from the Latin noun “cassis”, which means wearer of a helmet, in reference to the shape of the hind parts of the skull. The suggested English vernacular name is helmeted bat.
Geographic distribution: The type specimen of C. dimissus was collected by H. C. Robinson and E. Seimund in Kao Nawng, Bandon (currently within Tai Rom Yen National Park in Surat Thani province of Thailand) at 1,400 feet (= 436 m a.s.l.), in June 1913 (Robinson and Kloss 1915). However, the altitude associated with this specimen was reported by Thomas (1916) as 3,500 feet (= 1,067 m a.s.l.), which would correspond to near the summit of the Khao Nong mountain, where the collectors did not capture bats. As all known localities of Cassistrellus are located in the lowlands at elevation between 190 and 674 m a.s.l., these bats should be indeed regarded as lowland dwellers. The vast area covered by the few scattered records of Cassistrellus suggests that it should be widely distributed from the Isthmus of Kra into mainland Southeast Asia and the foothills of the Himalaya, i.e., across most of the Indo-Burma biodiversity hotspot (Myers et al. 2000a). All capture sites were situated in hilly terrain with mixed deciduous or dipterocarp forests traversed by large rivers.
Fig. 4. Lateral view of skull of A) Cassistrellus yokdonensis sp. nov. (holotype, ROM 107751) and B) C. dimissus(holotype, BM(NH) 16.4.21.1.). The scale bar at the bottom represents 5 mm.
Fig. 4. Lateral view of skull of C) C. dimissus from Laos (MHNG 1926.053), and D) C. dimissusfrom Nepal (UMMZ 172218). The scale bar at the bottom represents 5 mm.
Fig. 4. Lateral view of skull of A) Cassistrellus yokdonensis sp. nov. (holotype, ROM 107751), B) C. dimissus (holotype, BM(NH) 16.4.21.1.), C) C. dimissus from Laos (MHNG 1926.053), and D) C. dimissus from Nepal (UMMZ 172218).
Holotype: Male ROM 107751 (field number 42734) collected on 6 June 1997 by B. K. Lim and M. Theberge. Preserved as a skin, skull, and partial skeleton. Epiphyses almost completely fused, indicating that this bat was a subadult.
Type locality: Vietnam: Dak Lak province; Yok Don National Park, Dak Ken River (tributary of the Serepok River), base of Yok Mt. ..., at 194 m a.s.l. in dry, open dipterocarp forest.
Diagnosis: Cassistrellus yokdonensis sp. nov. is a medium-sized vespertilionid bat (body mass about 15 g) characterized by pointed, narrow wings similar in shape to those of Nyctalus species, but not as narrow. The fur is sparse with short hairs, clove brown (Ridgway 1912) dorsally, lighter beige ventrally, and cream colored at the throat. There is no glossy tinge to the fur. The color of the wings and other skin parts is blackish brown. The wing membranes attach to the distal end of the metatarsus. The calcar extends less than halfway to the tail and has no visible lobe.
Although the dental formula is identical to that of Eptesicus species, the skull possesses a pair of deep and well-defined basisphenoid pits and prominent preorbital processes that are otherwise absent from the latter genus. C. yokdonensis sp. nov. is morphologically similar to C. dimissus, but—in spite of the fact that the known individuals are not fully grown adults—is substantially larger, both externally (e.g., FA 47 mm versus 39–42 mm; Table 1), and cranially (e.g., maxillary toothrow length over 6.5 mm versus less than 6.4 mm). C. yokdonensis sp. nov. has also a much stronger dentition in general, and subequal small lower premolars, longer more curved upper canines, and procumbent upper incisors compared to its congener. Genetically, C. yokdonensis sp. nov. has unique mitochondrial (Cytb, Co1) and nuclear (Rag2) sequences compared to C. dimissus from Laos.
Etymology: We propose the name C. yokdonensis after the national park where it has been found, in recognition of the importance of protected areas in conserving species and their habitats.
Geographic distribution: Currently known only from Yok Don National Park, Dak Lak Province of Vietnam. The 2 specimens were caught shortly after 2100 h as they flew into large (30 m long by 10 m high) canopy nets deployed in a dry, open dipterocarp forest of lowland regions. Other species of mammals caught in the same nets included Taphozous, Pipistrellus, Murina, Cynopterus, Megaerops, and Rhinolophus bats and several Hylopetes flying squirrels. In addition, a paratype specimen of a new species of parachute gecko (Ptychozoon trinitaterra) was caught in this net (Brown 1999).
Manuel Ruedi, Judith L. Eger, Burton K. Lim and Gábor Csorba. 2017. A New Genus and Species of Vespertilionid Bat from the Indomalayan Region. Journal of Mammalogy. gyx156. DOI: 10.1093/jmammal/gyx156
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شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
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تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
The areolate Oriental family Heteropterygidae Kirby, 1893 is critically reviewed and the results of the present study contradict the arrangement suggested by Zompro (2004), but in most aspects agree with a molecular study presented by Whiting et al (2003) and a phylogenetic study presented by Bradler (2009). The family is critically discussed and new hypotheses are presented for the phylogeny and intra-familiar relationships, placing the subfamily Dataminae Rehn & Rehn, 1939 as the basalmost clade of Heteropterygidae. The subfamilies Obriminae Brunner v. Wattenwyl, 1893 and Heteropteryginae Kirby, 1893 together represent the sister-group of Dataminae. Arguments and a tree are presented to support this hypothesis. New diagnoses and lists of genera are provided for all three subfamilies contained in Heteropterygidae, along with keys to distinguish between them.
The subfamily Obriminae is critically reviewed and the distinction between the three tribes Obrimini Brunner v. Wattenwyl, 1893, Eubulidini Zompro, 2004 and Miroceramiini Zompro, 2004 introduced by Zompro (2004) is shown to be poorly supported. While Obrimini sensuZompro, 2004 is generally accepted (but now also contains genera that were placed in Eubulidini or Miroceramiini by Zompro (2004)), the tribes Eubulidini and Miroceramiini are not supported. A new arrangement is introduced, which is based on morphological characters neglected or overlooked by Zompro (2004) but were partly discussed by Bradler (2009). The genus Mearnsiana Rehn & Rehn, 1939 is removed from Miroceramiini and transferred to Obrimini. The genera EubulidesStål, 1877, Heterocopus Redtenbacher, 1906, TheramenesStål, 1875 and Stenobrimus Redtenbacher, 1906 are removed from Eubulidini and also transferred to Obrimini. Consequently, Eubulidini is synonymised with Obrimini (n. syn.). Miroceramiini is a monotypical tribe and only includes the Wallacean genus Miroceramia Günther, 1934. The new tribe Tisamenini n. trib. is established for the three basal genera TisamenusStål, 1875, IlocanoRehn & Rehn, 1939 and Hoploclonia Stål, 1875 all of which were placed in Eubulidini by Zompro (2004). The latter genus differs from the other two genera by the morphology of the female genitalia, which is unique amongst the entire family. Three generic groups are recognized within Obrimini, the Obrimus-group, Stenobrimus-group and Theramenes-group. Keys are presented to distinguish between the three tribes now contained in the Obriminae, i.e. Obrimini, Tisamenini n. trib. and Miroceramiini. The genus Hennobrimus Conle, 2006 is synonymised with Mearnsiana Rehn & Rehn, 1939, based on the fact that the type-species of both genera are conspecific (n. syn.). Hennobrimus hennemanni Conle, 2006, the type-species of Hennobrimus, and Trachyaretaon manobo Lit & Eusebio, 2005 are synonymised with Mearnsiana bullosa Rehn & Rehn, 1939, the type-species of Mearnsiana (n. syn.).Theramenes dromedarius Stål, 1877 from the Philippines is removed from synonymy with the Wallacean Theramenes olivaceus (Westwood, 1859) and re-established as a valid species (rev. stat.).
The subfamily Heteropteryginae Kirby, 1896 is revised at the species-level and a new diagnosis is presented. Keys to the two genera and all 16 known species are provided along with new descriptions, differential diagnoses, lists of examined material, detailed information on the known distributions, measurements and illustrations of the insects and eggs. The intra-subfamiliar and intra-generic relationships are discussed and a cladogram is presented. Heteropteryginae contains two genera: Heteropteryx Gray, 1835 (Type-species: Phasma dilatatum Parkinson, 1798) and Haaniella Kirby, 1896 (Type-species: Phasma (Heteropteryx) muelleri de Haan, 1842). The distribution of this subfamily is restricted to Sundaland with the exception of a single species that is found in Vietnam. All other species are distributed in Borneo, Sumatra, the Mentawai Islands, Singapore, Peninsular Malaysia and Thailand. Heteropteryginae contains the largest and most striking members of the entire family Heteropteryginae, some of which are amongst the heaviest insects known. The subfamily is characterized by apomorphies such as the presence of wings, having a tympanal area (= stridulatory organ) in the basal portion of the alae, straight profemora, strongly shortened tarsi, lack of rough sensory-areas on the prosternum and typically X-shaped micropylar plate of the eggs. The sister-group of Heteropteryginae is represented by the Obriminae, with which it shares a beak-like secondary ovipositor in the females and presence of a medio-apical spine on the area apicalis. Both features are synapomorphies of Heteropteryginae + Obriminae.
The genus Haaniella Kirby, 1904 contains 16 known species, five of which are newly described herein. The genus Miniopteryx Zompro, 2004 (Type-species: Haaniella parva Günther, 1944) is synonymised with Haaniella on the basis that the distinguishing feature mentioned in the original description is a character that is frequently found throughout the genus (n. syn.). The type-species H. parva Günther, 1944 is automatically retransferred to Haaniella (rev. stat.). Haaniella aculeata n. sp. from western Sumatra is described from the male. Haaniella macroptera n. sp. from Singapore and the Johor state in southern Peninsular Malaysia is described from both sexes and the eggs. Haaniella gintingi n. sp. from Central Sumatra is described from both sexes and the eggs and Haaniella kerincia n. sp. from Western Sumatra is described from the insects only, the eggs being still unknown. One new species, Haaniella gorochovi n. sp., is the only representative of the genus and subfamily Heteropteryginae known from Vietnam and both sexes as well as the eggs are described. Haaniella erringtoniae (Redtenbacher, 1906) is endemic in Peninsular Malaysia, here removed from synonymy with H. muelleri(de Haan, 1842) and re-established as a valid species (rev. stat.). The Sumatran Haaniellaglaber(Redtenbacher, 1906) is removed from synonymy with H. muelleri (Haan, 1842) and re-established as a valid species (rev. stat.). Leocrates glaber Redtenbacher, 1906 and Haaniella muelleri simplex Günther, 1944 are removed from synonymy with H. muelleri (Haan, 1842) (rev. stat.) and synonymised with H. glaber.Haaniellamecheli(Redtenbacher, 1906) andH. rosenbergii(Kaup, 1871) are removed from synonymy with H. muelleri (Haan, 1842) and re-established as valid species (rev. stat.). Haaniella erringtoniae novaeguineaeGünther, 1934 and Haaniella muelleri var. b. (Haan, 1842) are synonymized with H. rosenbergii (Kaup, 1871) (n. syn.). The type-species Haaniella muelleri (Haan, 1842) is shown to be a fairly rare species that is restricted to Sumatra. All subsequent records of H. muelleri from outside Sumatra and references to captive breeding of stock originating from Peninsular Malaysia in Europe relate to H. erringtoniae (Redtenbacher, 1906). The previously unknown males and eggs of H. rosenbergii (Kaup, 1871) as well as the previously unknown females and eggs of H. parva Günther, 1944 are described and illustrated for the first time. Based on morphological characters of the insects and eggs three distinct species-groups are recognized within Haaniella. The muelleri species-group contains nine species that are distributed throughout Sumatra, the Mentawei Islands, Singapore and Peninsular Malaysia. These are characterized by the smooth ventral surface of the meso- and metafemora and lemon-shaped eggs which entirely lack the setae seen in the two other species-groups. The grayii species-group comprises four species, two of which are endemic in Borneo, one endemic in Sumatra and the fourth species being the only known representative of the subfamily in Vietnam. These species are characteristic for the prominent pair of spines on the abdominal tergites II–IV of males and long apically multidentate epiproct of females. The echinata species-group contains three exceptionally Bornean species, which are characterized by the long and apically pointed subgenital plate of females, which clearly projects beyond the epiproct, as well as the sub-basal lateral tooth of the anal segment of males. The muelleri species-group is sister to the remainder two species-groups.
Heteropteryx Gray, 1853 is a monotypical genus and only contains the type-species H. dilatata (Parkinson, 1798), which is found throughout Peninsular Malaysia, Thailand, Sumatra and Northeastern Borneo. This genus differs from Haaniella by the strongly conically elevated head, which posteriorly projects over the anterior margin of the pronotum, females being bright green or yellow in colour with plain and translucent pink alae and having distinct spines on the abdominal tergites, and males having a strongly shortened mesothorax and dull pink alae.
Lectotypes are designated for Haaniella parva Günther, 1944, Heteropteryx echinata Redtenbacher, 1906, Heteropteryx saussurei Redtenbacher, 1906 and Heteropteryx scabraRedtenbacher, 1906 to guarantee stability of these names.
Information on the habitats, host-plants, biology, life cycle, parasitism and captive breeding of the species of Heteropteryginae is presented and a list summarising all taxonomic changes presented herein.
Keywords: Phasmatodea, Heteropterygidae, Heteropteryginae, Obriminae, Dataminae, Heteropteryx, Haaniella, taxonomic revision, classification, new tribe, new species, new subspecies, new synonyms, lectotypes, keys, differentiations, descriptions, illustrations, eggs
Frank H. Hennemann, Oskar V. Conle, Paul D. Brock and Francis Seow-Choen. 2016. Revision of the Oriental subfamily Heteropteryginae Kirby, 1896, with A Re-arrangement of the family Heteropterygidae and the Descriptions of Five New Species of Haaniella Kirby, 1904. (Phasmatodea: Areolatae: Heteropterygidae). Zootaxa. 4159(1); 1–219. DOI: 10.11646/zootaxa.4159.1.1
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
