Search This Blog

مشاركة مميزة

[Ornithology • 2017] Myzomela irianawidodoae • A Colourful New Species of Myzomela Honeyeater from Rote Island in eastern Indonesia ---ScRaBBlE

Myzomela irianawidodoae Prawiradilaga, Baveja, Suparno, Ashari, Ng, Gwee, Verbelen & Rheindt, 2017  photo:   Philippe Verbelen  e-journ...

Translate

Showing posts with label Tribe. Show all posts
Showing posts with label Tribe. Show all posts

Wednesday, March 20, 2019

[Ichthyology • 2017] Gymnogeophagus taroba • A New Species (Teleostei: Cichlidae) from the río Iguazú Basin, Misiones, Argentina ---ScRaBBlE


Gymnogeophagus taroba
 Casciotta, Almirón, Piálek & Říčan, 2017

HISTORIA NATURAL. Tercera Serie. 7(2)

Abstract

 The Gymnogeophagus setequedas group is based on our results composed of three endemic species of which one, Gymnogeophagus taroba sp. n., is described as a new species. The three species are diagnosable from each other and from other species of Gymnogeophagus by stable differences in several morphological characters among which the best are found in coloration patterns. Body and head shapes and meristic characters show lesser differentiation but several are also clearly diagnostic in the G. setequedas group. The G. setequedas group is strongly structured allopatrically. The prime candidates for this fragmentation and speciation are the origins of the waterfalls on the individual tributaries. The largest of the waterfalls, the famous Cataratas del Iguazú, with a height of 72 m, separate G. taroba sp. n. from its closest relatives G. che and G. setequedas. The original 28 m high Urugua-í falls separate G. che from G. setequedas. Gymnogeophagus setequedas is separated from G. che and G. taroba by large rapids (about 65m in total elevation above the río Paraná) and a former fall on the Acaray river and by the 45 m high Monday falls located a few km from the mouth of the Monday into the río Paraná just opposite the mouth of the Iguazú.

 Key words. Cichlid fishes, endemism, morphology, La Plata River basin, Paraná river basin


Figure 3- Gymnogeophagus taroba, coloration of live specimens immediatelly after capture.
 (A-B) IBIGEO-I 449, paratype specimens, 87.8 mm, 79.8 mm. (C-D) MLP uncat., Říčan et al., February 2014, data as holotype. 

Taxonomy
Family Cichlidae Bonaparte, 1835
Genus Gymnogeophagus Miranda Ribeiro, 1918

Gymnogeophagus taroba, new species (Figures 1-7; Table 1)

Gymnogeophagus aff. setequedas Casciotta et al., 2016
(photo of live holotype, first mention of the species)

Diagnosis. Gymnogeophagus taroba is distinguished from all species of Gymnogeophagus in the G. gymnogenys group by the possession of 23 to 25 E1 scales and the absence of a well developed adipose hump in adult males. Gymnogeophagus taroba is most easily distinguished from all species in the G. rhabdotus group (G. terrapurpura, G. rhabdotus, G. meridionalis, G. setequedas and G. che) by the pigmentation of the dorsal fin. ....

......

Etymology. The specific epithet taroba refers to Tarobá, a warrior, and refers to a legend of the Kaingang people; a noun in apposition. The Kaingang were the original first inhabitants of the present province of Misiones in Argentina and the southern Brazilian states of Paraná, Santa Catarina and Rio Grande do Sul and also the southeastern state of São Paulo. Their language and culture is quite distinct from the neighboring Guaraní. The Kaingang language is classified as a member of the Gê language family. The legend tells that at the beginning of time, the río Iguazú was inhabited by a huge and monstrous serpent called Mboi, a guardian god. The Kaingang sacrificed a beautiful young maiden every year to Mboi. When Tarobá meets Naipí, chosen for the sacrifice, he rebells against the elders of the tribe who refuse to release her. Tarobá and Naipí try to escape in a canoe by the river. Mboi becomes furious and brakes the course of the river forming the Iguazú falls, catching the lovers. Naipí is transformed into one of the rocks of the falls, perpetually punished by the turbulent waters, and Tarobá is turned into a palm tree on the bank of the waterfall. Mboi lives submerged in the Garganta del Diablo, from where he watches over the lovers, preventing them from joining again. However, on sunny days and as a bridge of love, the rainbow overcomes the power of Mboi by rejoining Naipí and Tarobá.


Jorge Casciotta, Adriana Almirón, Lubomir Piálek and Oldřich Říčan. 2017. Gymnogeophagus taroba (Teleostei: Cichlidae), A New Species from the río Iguazú Basin, Misiones, Argentina.  HISTORIA NATURAL. Tercera Serie. 7(2); 5-22.


Resumen. En base a nuestros resultados, el grupo Gymnogeophagus setequedas esta compuesto por tres especies endémicas, de las cuales, G. taroba sp. n., se describe como una nueva especie. Las tres especies son diagnosticables entre sí y de otras especies de Gymnogeophagus por diferencias estables en varios caracteres morfológicos, entre los cuales los mejores son los patrones de coloración. La forma del cuerpo, la cabeza y los caracteres merísticos muestran una menor diferenciación, pero algunos también son claramente diagnósticos en el grupo G. setequedas. El grupo G. setequedas está fuertemente estructurado alopátricamente. La causa principal para esta fragmentación y especiación es la formación de cascadas en los afluentes. La más grande de las cascadas, la famosa Catarata del Iguazú, con una altura de 72 m, separa G. taroba sp. n. de sus parientes más cercanos G. che y G. setequedas. El salto original del Urugua–í de 28 m de altura, separa G. che de G. setequedas. Gymnogeophagus setequedas está separada de G. che y G. taroba por grandes rápidos (unos 65 m de elevación total sobre el río Paraná) y un primer salto en el río Acaray y por otro salto de 45 m de altura en el río Monday localizado a unos pocos kilómetros de la desembocadura del Monday en el río Paraná justo enfrente de la desembocadura del Iguazú. 
Palabras clave. Cíclidos, endemismo, morfología, Cuenca del Plata, Cuenca del río Paraná

---------------------------------------------------------------
روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Herpetology • 2018] Rana lenca • An Integrative Assessment of the Taxonomic Status of Putative Hybrid Leopard Frogs (Anura: Ranidae) from the Chortís Highlands of Central America, with Description of A New Species ---ScRaBBlE


Rana lenca
 Luque-Montes, Austin, Weinfurther, Wilson, Hofmann & Townsend, 2018


Integrative taxonomy seeks to approach the complex topic of species diagnosis using independent, complementary lines of evidence. Despite their ubiquity throughout North and Central America, taxonomy of the American leopard frogs (Anura: Ranidae: Rana: subgenus Pantherana) remains largely unresolved, and this is arguably nowhere truer than in the Central American country of Honduras, where there are two nominal species, the taxonomy of which remains unresolved. Leopard frogs from several mountainous areas along the continental divide in Honduras have previously been considered putative hybrids between Rana brownorum and R. cf. forreri, as opposed to two alternate hypotheses: one that they represent a high-altitude eco-morph of a single widespread species that included both lowland forms, or a second that there is an undescribed highland species distinct from either of the recognized lowland forms. We examine this set of hypotheses using three independent lines of evidence. First, we used species distribution modelling to examine potential geographic isolation of the highland form and the two putative parental lowland species, and found strong ecological separation between the highland and lowland forms. Second, mitochondrial and nuclear DNA supports the distinction of the highland form from both putative parental species, with mtDNA data refuting the hypothesis that representatives of either species may represent a matrilineal founder. Morphologically, the highland form is significantly smaller than, and otherwise readily differentiated from, both R. brownorum and R. cf. forreri, as well as all other Rana found in Honduras and adjacent areas. As a result, we formally describe the highland leopard frog as a new species.

Key words: Amphibia, integrative taxonomy, mitochondrial DNA, Pantherana, phylogeny, rhodopsin, species distribution modelling




Rana lenca sp. nov.

Common English name. Lenca Leopard Frog
Common Spanish name. La Rana Lenca

Etymology. The name lenca is given in honour of the indigenous Lenca people, the traditional inhabitants of the mountainous region of south-western Honduras. 

Paratypes of Rana lenca from Reserva Biologica Cerro Uyuca, 1,640 m elevation, Departamento de Francisco Morazan,Honduras:
(1) adult female paratype (UF 166642; 64.3 mm SL); (2) Subadult female paratype (UF 166643) and tadpole (UF 166637).
Photos by Jason M. Butler. 

Unvouchered examples of Rana lenca; (1) adult male from the type locality; (2) adult female from the type locality; (3) adult male from the Reserva Biologica Cerro Uyuca; (4) adult female from the Reserva Biologica Cerro Uyuca. 

Unvouchered examples of Rana lenca;
 (1) adult male from the type locality; (2) adult female from the type locality; (3) adult male from the Reserva Biologica Cerro Uyuca; (4) adult female from the Reserva Biologica Cerro Uyuca; (5) adult male at edge of pond at type locality; (6) adult female floating amongst Pinus oocarpa needles in a spring-fed pool at Reserva Biologica Cerro Uyuca. 

Type locality of Rana lenca; San Pedro La Loma, 2010 m elevation, Depto. Intibuca, Honduras. in January 2008 (top) and during drought conditions in May 2015 (bottom). 


Ileana Luque-Montes, James D. Austin, Kayla D. Weinfurther, Larry David Wilson, Erich P. Hofmann and Josiah H. Townsend. 2018. An Integrative Assessment of the Taxonomic Status of Putative Hybrid Leopard Frogs (Anura: Ranidae) from the Chortís Highlands of Central America, with Description of A New Species.  Systematics and Biodiversity. In Press.   DOI  10.1080/14772000.2017.1415232 
---------------------------------------------------------------
روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Mammalogy • 2018] Uncovering the Diversity of Dog-faced Bats from the Genus Cynomops (Chiroptera: Molossidae), with the Redescription of C. milleri and the Description of Two New Species; Cynomops freemani & C. tonkigui ---ScRaBBlE


Cynomops freemani 
Moras, Gregorin, Sattler & Tavares, 2018


Abstract
Until recently the genus Cynomops (Chiroptera: Molossidae) comprised six species of fast flying, aerial insectivorous bats distributed throughout the Neotropics. However, our revisionary study have shown that the diversity of this taxon was underestimated as molecular and morphological data supported the recognition of C. milleri as a full species and revealed two previously unrecognized, small species of Cynomops. These newly recognized taxa are Cynomops freemani sp. nov. from the Canal Zone region, Panama, sister taxon to C. mexicanus, and Cynomops tonkigui sp. nov. from the eastern Andes of Ecuador and Colombia, sister taxon of C. milleri. We also describe the calls of Cynomops freemani sp. nov. that may be useful for the recognition of these species during field inventories based on ultrasonic recording techniques. We elevate the currently known diversity of Cynomops to eight species, as we also revalidate C. milleri providing an emended diagnosis and a redescription of this taxon.

Keywords: Free-tailed bats, taxonomy, acoustic identification, Cynomops freemani sp. nov, Cynomops tonkigui sp. nov

Systematics
Family Molossidae Gervais 1856
Genus Cynomops Thomas 1920

Cynomops milleri (Osgood, 1914)
Miller’s Dog-faced Bat




two Freeman’s Dog-Faced bats Cynomops freemani discovered in Gamboa, Panama.
Photo: Elias Bader

Cynomops freemani, new species 
Freeman’s Dog-faced Bat

ETYMOLOGY: The name “freemani” is in honor to Dr. Patricia W. Freeman in recognition of her influential contributions to the study of the morphology, systematics, and evolutionary relationships of molossid bats (e.g. Freeman, 1981).

DISTRIBUTION: Cynomops freemani is known from lowlands (23–53 m a.s.l.) of the Pacific coast of the Canal Zone region in Panama (Fig. 3).


 Waorani dog-faced bat Cynomops tonkigui 
Photo: Diego Tirira

Cynomops tonkigui, new species 
Waorani Dog-faced Bat

ETYMOLOGY: The name “tonkigui” is used as a noun in apposition, and means "bat" in the Waorani language. The name honors the Waorani people who inhabit the type locality (Tirira, 2012)

DISTRIBUTION: Cynomops tonkigui is known from the lowlands (195–529 m a.s.l.) of northeastern South America, and the eastern slopes of the Andes in Ecuador and Colombia (Fig. 3).


 Ligiane M. Moras, Renato Gregorin, Thomas Sattler and Valéria da C. Tavares. 2018. Uncovering the Diversity of Dog-faced Bats from the Genus Cynomops (Chiroptera: Molossidae), with the Redescription of C. milleri and the Description of Two New Species. Mammalian Biology. In Press. DOI: 10.1016/j.mambio.2017.12.005 

   

Two New Dog-Faced Bat Species Discovered in Panama and Ecuador newsdesk.si.edu/releases/two-new-dog-faced-bat-species-discovered-panama-and-ecuador
More sky puppies! Scientists discover two new species of dog-faced bat insider.si.edu/2018/01/sky-puppies-scientists-discover-two-new-species-dog-faced-bat 

---------------------------------------------------------------
روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Ichthyology • 2018] Spectracanthicus javae • A New Species of the Genus Spectracanthicus (Loricariidae, Hypostominae, Ancistrini) from the Rio Javaיs (Rio Araguaia Basin), with A Description of Gross Brain Morphology ---ScRaBBlE


Spectracanthicus javae 
Chamon, Pereira, Mendonca & Akama, 2018

  DOI:  10.1111/jfb.13526  

Abstract
A new species of Spectracanthicus is described from the Rio Javaés, Rio Araguaia basin. The new species is distinguished from its congeners (except Spectracanthicus immaculatus) by colour pattern: body dark grey to dark brown without dots or blotches (v. body colour with yellowish small dots in Spectracanthicus murinusSpectracanthicus punctatissimus and Spectracanthicus tocantinensis and large white dots in Spectracanthicus zuanoni). It can be further distinguished from S. immaculatus by having thicker and less numerous teeth, with up to eight premaxillary and 20 dentary teeth (v. teeth thinner and more numerous with up to 22 premaxillary and 30 dentary teeth); dorsal and caudal fins without curved spines (v. dorsal and caudal fins with curved spines). Other osteological characters can also diagnose the new species from its congeners. In addition, a gross brain description and brief comments on the new species' ecological habitat are given.


Figure 2. Colour in life of Spectracanthicus javae.

Spectracanthicus javae sp. nov.

Etymology: The specific epithet (a noun in apposition) is in reference to the Javaé people, an indigenous branch of the Karajá people in Brazil who inhabit the Ilha do Bananal and confer the name of the river (Rio Javaés), where the new species was reported.



C. C. Chamon, T. N. A. Pereira, M. B. Mendonca and A. Akama. 2018.  New Species of the Genus Spectracanthicus (Loricariidae, Hypostominae, Ancistrini) from the Rio Javaיs (Rio Araguaia Basin), with A Description of Gross Brain Morphology. Journal of Fish Biology.   DOI:  10.1111/jfb.13526 

Carine C. Chamon and Lúcia H. Rapp Py-Daniel. 2014. Taxonomic revision of Spectracanthicus Nijssen & Isbrücker (Loricariidae: Hypostominae: Ancistrini), with description of three new species. Neotropical Ichthyology. 12(1):1-25. 


---------------------------------------------------------------
روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Entomology • 2016] Revision of the Oriental subfamily Heteropteryginae Kirby, 1896 (Phasmatodea: Areolatae: Heteropterygidae), with A Re-arrangement of the family Heteropterygidae and the Descriptions of Five New Species of Haaniella Kirby, 1904 ---ScRaBBlE


Heteropteryx dilatata  (Parkinson, 1798)


Abstract  

The areolate Oriental family Heteropterygidae Kirby, 1893 is critically reviewed and the results of the present study contradict the arrangement suggested by Zompro (2004), but in most aspects agree with a molecular study presented by Whiting et al (2003) and a phylogenetic study presented by Bradler (2009). The family is critically discussed and new hypotheses are presented for the phylogeny and intra-familiar relationships, placing the subfamily Dataminae Rehn & Rehn, 1939 as the basalmost clade of Heteropterygidae. The subfamilies Obriminae Brunner v. Wattenwyl, 1893 and Heteropteryginae Kirby, 1893 together represent the sister-group of Dataminae. Arguments and a tree are presented to support this hypothesis. New diagnoses and lists of genera are provided for all three subfamilies contained in Heteropterygidae, along with keys to distinguish between them.

        The subfamily Obriminae is critically reviewed and the distinction between the three tribes Obrimini Brunner v. Wattenwyl, 1893, Eubulidini Zompro, 2004 and Miroceramiini Zompro, 2004 introduced by Zompro (2004) is shown to be poorly supported. While Obrimini sensu Zompro, 2004 is generally accepted (but now also contains genera that were placed in Eubulidini or Miroceramiini by Zompro (2004)), the tribes Eubulidini and Miroceramiini are not supported. A new arrangement is introduced, which is based on morphological characters neglected or overlooked by Zompro (2004) but were partly discussed by Bradler (2009). The genus Mearnsiana Rehn & Rehn, 1939 is removed from Miroceramiini and transferred to Obrimini. The genera Eubulides Stål, 1877Heterocopus Redtenbacher, 1906Theramenes Stål, 1875 and Stenobrimus Redtenbacher, 1906 are removed from Eubulidini and also transferred to Obrimini. Consequently, Eubulidini is synonymised with Obrimini (n. syn.). Miroceramiini is a monotypical tribe and only includes the Wallacean genus Miroceramia Günther, 1934. The new tribe Tisamenini n. trib. is established for the three basal genera Tisamenus Stål, 1875Ilocano Rehn & Rehn, 1939 and Hoploclonia Stål, 1875 all of which were placed in Eubulidini by Zompro (2004). The latter genus differs from the other two genera by the morphology of the female genitalia, which is unique amongst the entire family. Three generic groups are recognized within Obrimini, the Obrimus-group, Stenobrimus-group and Theramenes-group. Keys are presented to distinguish between the three tribes now contained in the Obriminae, i.e. Obrimini, Tisamenini n. trib. and Miroceramiini. The genus Hennobrimus Conle, 2006 is synonymised with Mearnsiana Rehn & Rehn, 1939, based on the fact that the type-species of both genera are conspecific (n. syn.). Hennobrimus hennemanni Conle, 2006, the type-species of Hennobrimus, and Trachyaretaon manobo Lit & Eusebio, 2005 are synonymised with Mearnsiana bullosa Rehn & Rehn, 1939, the type-species of Mearnsiana (n. syn.). Theramenes dromedarius Stål, 1877 from the Philippines is removed from synonymy with the Wallacean Theramenes olivaceus (Westwood, 1859) and re-established as a valid species (rev. stat.).

        The subfamily Heteropteryginae Kirby, 1896 is revised at the species-level and a new diagnosis is presented. Keys to the two genera and all 16 known species are provided along with new descriptions, differential diagnoses, lists of examined material, detailed information on the known distributions, measurements and illustrations of the insects and eggs. The intra-subfamiliar and intra-generic relationships are discussed and a cladogram is presented. Heteropteryginae contains two genera: Heteropteryx Gray, 1835 (Type-species: Phasma dilatatum Parkinson, 1798) and Haaniella Kirby, 1896 (Type-species: Phasma (Heteropteryx) muelleri de Haan, 1842). The distribution of this subfamily is restricted to Sundaland with the exception of a single species that is found in Vietnam. All other species are distributed in Borneo, Sumatra, the Mentawai Islands, Singapore, Peninsular Malaysia and Thailand. Heteropteryginae contains the largest and most striking members of the entire family Heteropteryginae, some of which are amongst the heaviest insects known. The subfamily is characterized by apomorphies such as the presence of wings, having a tympanal area (= stridulatory organ) in the basal portion of the alae, straight profemora, strongly shortened tarsi, lack of rough sensory-areas on the prosternum and typically X-shaped micropylar plate of the eggs. The sister-group of Heteropteryginae is represented by the Obriminae, with which it shares a beak-like secondary ovipositor in the females and presence of a medio-apical spine on the area apicalis. Both features are synapomorphies of Heteropteryginae + Obriminae.

        The genus Haaniella Kirby, 1904 contains 16 known species, five of which are newly described herein. The genus Miniopteryx Zompro, 2004 (Type-species: Haaniella parva Günther, 1944) is synonymised with Haaniella on the basis that the distinguishing feature mentioned in the original description is a character that is frequently found throughout the genus (n. syn.). The type-species H. parva Günther, 1944 is automatically retransferred to Haaniella (rev. stat.). Haaniella aculeata n. sp. from western Sumatra is described from the male. Haaniella macroptera n. sp. from Singapore and the Johor state in southern Peninsular Malaysia is described from both sexes and the eggs. Haaniella gintingi n. sp. from Central Sumatra is described from both sexes and the eggs and Haaniella kerincia n. sp. from Western Sumatra is described from the insects only, the eggs being still unknown. One new species, Haaniella gorochovi n. sp., is the only representative of the genus and subfamily Heteropteryginae known from Vietnam and both sexes as well as the eggs are described. Haaniella erringtoniae (Redtenbacher, 1906) is endemic in Peninsular Malaysia, here removed from synonymy with H. muelleri (de Haan, 1842) and re-established as a valid species (rev. stat.). The Sumatran Haaniella glaber (Redtenbacher, 1906) is removed from synonymy with H. muelleri (Haan, 1842) and re-established as a valid species (rev. stat.). Leocrates glaber Redtenbacher, 1906 and Haaniella muelleri simplex Günther, 1944 are removed from synonymy with H. muelleri (Haan, 1842) (rev. stat.) and synonymised with H. glaber. Haaniella mecheli (Redtenbacher, 1906) and H. rosenbergii (Kaup, 1871) are removed from synonymy with H. muelleri (Haan, 1842) and re-established as valid species (rev. stat.). Haaniella erringtoniae novaeguineae Günther, 1934 and Haaniella muelleri var. b. (Haan, 1842) are synonymized with H. rosenbergii (Kaup, 1871) (n. syn.). The type-species Haaniella muelleri (Haan, 1842) is shown to be a fairly rare species that is restricted to Sumatra. All subsequent records of H. muelleri from outside Sumatra and references to captive breeding of stock originating from Peninsular Malaysia in Europe relate to H. erringtoniae (Redtenbacher, 1906). The previously unknown males and eggs of H. rosenbergii (Kaup, 1871) as well as the previously unknown females and eggs of H. parva Günther, 1944 are described and illustrated for the first time. Based on morphological characters of the insects and eggs three distinct species-groups are recognized within Haaniella. The muelleri species-group contains nine species that are distributed throughout Sumatra, the Mentawei Islands, Singapore and Peninsular Malaysia. These are characterized by the smooth ventral surface of the meso- and metafemora and lemon-shaped eggs which entirely lack the setae seen in the two other species-groups. The grayii species-group comprises four species, two of which are endemic in Borneo, one endemic in Sumatra and the fourth species being the only known representative of the subfamily in Vietnam. These species are characteristic for the prominent pair of spines on the abdominal tergites II–IV of males and long apically multidentate epiproct of females. The echinata species-group contains three exceptionally Bornean species, which are characterized by the long and apically pointed subgenital plate of females, which clearly projects beyond the epiproct, as well as the sub-basal lateral tooth of the anal segment of males. The muelleri species-group is sister to the remainder two species-groups.

        Heteropteryx Gray, 1853 is a monotypical genus and only contains the type-species H. dilatata (Parkinson, 1798), which is found throughout Peninsular Malaysia, Thailand, Sumatra and Northeastern Borneo. This genus differs from Haaniella by the strongly conically elevated head, which posteriorly projects over the anterior margin of the pronotum, females being bright green or yellow in colour with plain and translucent pink alae and having distinct spines on the abdominal tergites, and males having a strongly shortened mesothorax and dull pink alae. 

Lectotypes are designated for Haaniella parva Günther, 1944, Heteropteryx echinata Redtenbacher, 1906, Heteropteryx saussurei Redtenbacher, 1906 and Heteropteryx scabra Redtenbacher, 1906 to guarantee stability of these names.

        Information on the habitats, host-plants, biology, life cycle, parasitism and captive breeding of the species of Heteropteryginae is presented and a list summarising all taxonomic changes presented herein.

Keywords: Phasmatodea, Heteropterygidae, Heteropteryginae, Obriminae, Dataminae, HeteropteryxHaaniella, taxonomic revision, classification, new tribe, new species, new subspecies, new synonyms, lectotypes, keys, differentiations, descriptions, illustrations, eggs




Frank H. Hennemann, Oskar V. Conle, Paul D. Brock and Francis Seow-Choen. 2016. Revision of the Oriental subfamily Heteropteryginae Kirby, 1896, with A Re-arrangement of the family Heteropterygidae and the Descriptions of Five New Species of Haaniella Kirby, 1904. (Phasmatodea: Areolatae: Heteropterygidae).  Zootaxa. 4159(1); 1–219. DOI: 10.11646/zootaxa.4159.1.1

---------------------------------------------------------------
روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Nakonanectes bradti • A New Elasmosaurid (Sauropterygia, Plesiosauria) from the Bearpaw Shale (Late Cretaceous, Maastrichtian) of Montana Demonstrates Multiple Evolutionary Reductions of Neck Length within Elasmosauridae ---ScRaBBlE


Nakonanectes bradti 
Serratos, Druckenmiller, & Benson, 2017 


ABSTRACT
Plesiosauria is a diverse clade of marine reptiles that have been studied since the early 19th century. However, phylogenetic relationships within the group have been contentious due to limited taxon sampling and a misunderstanding of how ontogeny and interspecific and intraspecific variation affect character states. This is particularly true for elasmosaurids, a Cretaceous clade of long-necked plesiosaurians. In 2010, a new, nearly complete skeleton, MOR 3072, was collected from the Late Cretaceous (Campanian–Maastrichtian) Bearpaw Shale of northeast Montana. The specimen provides detailed morphological information rarely observed within Elasmosauridae, including a complete skull, the anterior 23 cervical vertebrae, a partial dorsal and caudal vertebral column, incomplete pectoral and pelvic girdles, elements of both fore- and hind limbs, ribs, and gastralia. Being early Maastrichtian in age, MOR 3072 is the stratigraphically youngest elasmosaurid known from the Western Interior Seaway and is recognized as a new genus and speciesNakonanectes bradti. We present a description of Nakonanectes bradti and conduct an extended phylogenetic analysis of Elasmosauridae. N. bradti is found to be deeply nested within the clade of large-bodied, long-necked, Western Interior Styxosaurinae. However, MOR 3072 is one of the smallest adult elasmosaurids ever recovered (5.1–5.6 m) and exhibits a reduced neck length compared with other North American elasmosaurids, resulting from a reduction in both centrum length and number of cervical vertebrae (39–42 were originally present). These features are convergent with the Southern Hemisphere clade of short-necked Maastrichtian elasmosaurids, Aristonectinae, and demonstrate multiple origins of short-necked body proportions from longer-necked ancestors within Elasmosauridae.

The newly named short-necked elasmosaur, Nakonanectes bradti, swims through an ancient sea in this artist’s reconstruction.
Illustration by James Havens, @alaskapaleoproject

 

Laser Scan of Cranium of MOR 3072, holotype specimen of Nakonanectes bradti, gen. et sp. nov.   

SYSTEMATIC PALEONTOLOGY
DIAPSIDA Osborn, 1903
SAUROPTERYGIA Owen, 1860

PLESIOSAURIA de Blainville, 1835
XENOPSARIA Benson and Druckenmiller, 2014

ELASMOSAURIDAE Cope, 1869
STYXOSAURINAE Otero, 2016

Phylogenetic Definition:— The clade including all taxa more closely related to Styxosaurus snowii than to Aristonectes parvidens (revised from Otero, 2016).


NAKONANECTES, gen. nov.
Type and Only Species:— Nakonanectes bradti.
Horizon:— Bearpaw Shale, lower Maastrichtian, Upper Cretaceous.


NAKONANECTES BRADTI, gen. et sp. nov. 
Holotype and Only Specimen:— MOR 3072, including a complete skull, articulated anterior 23 cervical vertebrae, dorsal and anterior caudal vertebrae, partial coracoids, much of the forelimbs including partial humeri, complete epipodial and mesopodial rows and associated phalanges, both ilia and portions of the pubis and ischium, portions of the hind limb including one complete and one partial femur with portions of the epipodial row, and numerous ribs and gastralia.

Etymology:— The generic name is derived from Nakona, the name for native Assiniboine people of northeastern Montana, which means ‘the friendly people.’ The specific name honors David Bradt of Florence, Montana, who found the specimen.

Nakonanectes bradti Serratos, Druckenmiller, & Benson, 2017



Serratos, D. J., P. Druckenmiller, and R. B. J. Benson. 2017. A New Elasmosaurid (Sauropterygia, Plesiosauria) from the Bearpaw Shale (Late Cretaceous, Maastrichtian) of Montana Demonstrates Multiple Evolutionary Reductions of Neck Length within Elasmosauridae. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1278608.

Alaska scientists help name new elasmosaur species from Montana - UAF News and Information http://news.uaf.edu/alaska-scientists-help-name-new-elasmosaur-species-from-montana/


---------------------------------------------------------------
روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

المشاركات الشائعة