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[Ornithology • 2017] Myzomela irianawidodoae • A Colourful New Species of Myzomela Honeyeater from Rote Island in eastern Indonesia ---ScRaBBlE

Myzomela irianawidodoae Prawiradilaga, Baveja, Suparno, Ashari, Ng, Gwee, Verbelen & Rheindt, 2017  photo:   Philippe Verbelen  e-journ...

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Showing posts with label Philippines. Show all posts
Showing posts with label Philippines. Show all posts

Wednesday, March 20, 2019

[Entomology • 2018] Colilodion colongi • A New Species of the Genus Colilodion Besuchet, 1991 (Coleoptera: Staphylinidae: Pselaphinae) from Mindanao, the Philippines ---ScRaBBlE


Colilodion colongi 
Hlaváč, Vondráček & Mohagan, 2018


Abstract

Colilodion colongi sp. nov. of the clavigerite ant-loving beetle from Mindanao, the Philippines is described, illustrated, and distinguished from related species. The distribution map for all known species is given.

Keywords: Coleoptera, Staphylinidae, Pselaphinae, Clavigeritae, Colilodionini, Colilodion, new species, Mindanao, the Philippines




Peter Hlaváč, Dominik Vondráček and Alma B. Mohagan. 2018. A New Species of the Genus Colilodion Besuchet, 1991 (Coleoptera: Staphylinidae: Pselaphinae) from Mindanao, the Philippines. Zootaxa. 4370(5); 562-568. DOI:  10.11646/zootaxa.4370.5.7

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Herpetology • 2018] Calliophis salitan • A New Species of Long-glanded Coralsnake of the Genus Calliophis (Squamata: Elapidae) from Dinagat Island, Philippines, with Notes on the Biogeography and Species Diversity of Philippine Calliophis and Hemibungarus ---ScRaBBlE


Calliophis salitan
Brown, Smart, Leviton & Smith, 2018


Abstract 
We describe a new species of coralsnake from northern Dinagat Island, southeastern Philippines. The discovery of this new species comes as a surprise because it is phenotypically distinct from all other Philippine coralsnakes and has a close phylogenetic affinity to the blue coralsnakes of the Sunda Shelf. The new species is distinguished from all Philippine and other Southeast Asian taxa by its large body size and ventral scale counts; its black head and neck; an alternating, broadly banded color pattern of black and off-white; and a bright orange tail. We use DNA sequence data to investigate the phylogenetic placement of the new species and that of several other populations of Philippine coralsnakes with respect to other Southeast Asian and Australasian elapids. Our results corroborate the uniqueness of the new species with respect to all other Philippine and Sundaic taxa, including the species most closely related to it: Calliophis bivirgatus, C. bilineatus, C. philippinus, and C. suluensis. We summarize phylogenetic, biogeographic, and phenotypic character data that substantiate the elevation of Philippine species of Calliophis (formerly considered subspecies of C. intestinalis: C. bilineatus, C. philippinus, and C. suluensis) and Hemibungarus (formerly subspecies of H. calligaster: H. calligaster, H. gemianulis, and H. mcclungi) to the level of full species. The allopatric distributions of these taxa emphasize the systematic and biogeographical significance of the newly discovered taxon: a poorly understood and independent colonization of the Philippine Archipelago by elapid snakes.

Keywords: Calliophis salitan, Caraga Region, Mindanao PAIC, new species, false coralsnakes




 Calliophis salitan adult male holotype (PNM 9844, formerly KU 310164; 997 mm total length [TL]), illustrated by E.F. Jones. 

Calliophis salitan sp. nov.

Etymology.— The specific epithet is a noun in apposition and is derived from the Tagalog (Filipino) term salitan, meaning‘‘alternating,’’in reference to the distinctive, alternating black and off-white banded color pattern characteristic of the new species.
Suggested English common name: Dinagat Island Banded Coralsnakes.

Habitat, distribution, and natural history.— The only known locality for Calliophis salitan is the low foothill of Mt. Cambinlia, Barangay Santiago, Municipality Loreto, Dinagat Island (Fig. 8), Philippines, where it was collected from an elevation of 195 m. This locality, dominated by selectively logged, regenerating second-growth forest, is situated at the northern end of the island, and has a tropical climate, with a mean annual precipitation of 3580 mm (recorded at Loreto town proper,,1 km from the type locality; higher precipitation values are possible at higher elevations on Mount Cambinlia). Precipitation is highest during December (~539 mm) and lowest in August (averaging 143 mm),and the mean annual temperature is 278C. The type specimen was collected between 1900 and 2200 h, and was actively crawling along a stream bed with reduced water flow between intermittent pools. The specimen was obtained at the end of July, toward the end of the dry season.


Rafe M. Brown, Utpal Smart, Alan E. Leviton, and Eric N. Smith. 2018. A New Species of Long-glanded Coralsnake of the Genus Calliophis (Squamata: Elapidae) from Dinagat Island, with Notes on the Biogeography and Species Diversity of Philippine Calliophis and HemibungarusHerpetologica. In-Press. DOI: 10.1655/Herpetologica-D-17-00008

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Entomology • 2017] Leaf Insects (Phasmida: Phylliidae) from Luzon, Philippines, with Descriptions of Four New Species, the New Genus Pseudomicrophyllium, and Redescription of Phyllium (Phyllium) geryon Gray, 1843 ---ScRaBBlE


 Pseudomicrophyllium faulkneri Cumming

in Cumming, Leong & Lohman, 2017. 

Abstract

Examination of unidentified Phylliidae specimens revealed a number of undescribed species from the island of Luzon, Philippines. Morphological and molecular study of specimens from the obscure phasmid genus Microphyllium Zompro, 2001, revealed a new species, which we describe as Microphyllium haskelli Cumming sp. nov.. It is here described and differentiated from the two other species in the genus, both currently only known from adults of a single sex. Pseudomicrophyllium Cumming gen. nov. is described as a new genus within Phylliidae with the type species Pseudomicrophyllium faulkneri Cumming gen. et sp. nov. as the sole known species in the genus. As is unfortunately often the case in the leaf-mimicking family Phylliidae, this new genus and species is only known from a single specimen. In addition to the new genus, two new Phyllium (Phyllium) species from the siccifolium species-group are named and described as Ph. (Ph.) antonkozlovi Cumming sp. nov. and Ph. (Ph.) bourquei Cumming & Le Tirant sp. nov.. In addition to the newly described species, Phyllium (Phyllium) geryon Gray, 1843 is redescribed from a nearly perfect specimen, completing some of the morphological knowledge gaps currently missing because of the severely damaged holotype specimen. A key to all known species of Phylliidae from Luzon is included. Holotype specimens for all four new species will be deposited in the National Museum of the Philippines type collection and paratype specimens will be deposited into the San Diego Natural History Museum collection or retained within the first author’s collection.

Keywords: Phasmatodea, DNA barcodes, Philippines, Luzon, new species, Phylliinae, holotype, Phyllium, Mountain Province, leaf insect

FIGURE 3. Microphyllium haskelli Cumming sp. nov., PT [Coll. RC 16-128]. A: dorsal view; B: ventral view.

Microphyllium haskelli Cumming, gen. nov.  

Etymology. This new species is a patronym named in honor of world famous forensic entomologist Dr. Neal H. Haskell of Rensselaer, Indiana, United States. Dr. Haskell has been an invaluable mentor to the first author as he works on his master’s degree.


FIGURE 8. Male Pseudomicrophyllium faulkneri Cumming gen. et sp. nov., HT [Coll.RC 16-129], A: dorsal view; B: ventral view

Pseudomicrophyllium Cumming gen. nov. 
Type Species: Pseudomicrophyllium faulkneri Cumming gen. et sp. nov. 

Etymology: The genus is named because it was thought for several years to be a new Microphyllium species because of its size and it was not until closer examination that it was revealed it to be a new genus evading detection right under the first author’s view because of those similarities. 

Diagnosis. ♂: Appearing closely related to Phyllium (Phyllium) with characteristics such as the antennae, thorax, but the body size and spination of the legs are reminiscent of Microphyllium. The combination of the below morphological features characterize males of the new genus and differentiate it from the other known phylliid genera.

Antennae: Long and filiform with long setae, similar to Phyllium (Phyllium).
Profemora: Completely lacking an exterior lobe, interior lobe greatly reduced, similar to Microphyllium.
Protibia: Lacking both exterior and interior lobes, similar to Microphyllium.
Thorax: Similar to Phyllium (Phyllium) with a distinct mesopraescutum.

The most evident feature that differentiates Pseudomicrophyllium gen. nov. from Microphyllium Zompro, 2001 are the long filiform antennae. Abdominal shape in the walking leaf family is a feature that has proven to be very variable in female Phyllium and more stable in male Phyllium. However not enough is known about the male variation of abdominal shape in the small phylliid genera of Luzon to use abdominal shape as a strong morphological feature in their differentiation and identification. Antennae structure is a feature that has proven much more stable throughout the family and this emphasis is reflected in the dichotomous key at the end of this work.


Pseudomicrophyllium faulkneri Cumming sp. nov.

Etymology. This new species is a patronym named in honor of David Faulkner of California, USA. Faulkner has been a mentor to the first author over the years and was the first to introduce him to forensic entomology. 

  
FIGURE 12. Phyllium (Phyllium) antonkozlovi Cumming sp. nov., HT, [Coll. RC 16-122], A: dorsal view; B: ventral view.

Phyllium (Phyllium) antonkozlovi Cumming sp. nov.

Etymology. This new species is a patronym named in honor of Anton Olegovich Kozlov, long time friend to the first author and well-known Russian field entomologist. 


FIGURE 14. Phyllium (Phyllium) bourquei Cumming & Le Tirant sp. nov. HT female [Coll. RC 17-203], A: dorsal view; B: ventral view; C: left foreleg.


Phyllium (Phyllium) bourquei Cumming & Le Tirant sp. nov.

 Etymology. This species is dedicated to Mr. Pierre Bourque. He was mayor of the City of Montreal from 1994 to 2001. Mr. Bourque was also one of the most innovative directors of the Montreal Botanical Garden from 1980 to 1994. Among other things, he was responsible for the creation of new greenhouses, the Floralies Internationales de Montréal, the Japanese Garden, the Chinese Garden, the Tree House, the Montreal Biodome and the Montreal Insectarium. Thanks to Mr. Bourque, the Montreal Botanical Garden has become the second largest in the world.


Checklist of Phylliidae species currently confirmed from Luzon, Philippines, and their recorded ranges. See figure 21 for an illustrated map of the distributions.


Phylliidae, Phylliinae, Phylliini:
 • Microphyllium Zompro, 2001
Microphyllium pusillulum (Rehn & Rehn, 1933)  [Central Luzon (Mountain* & Nueva Vizcaya Prov.)]
Microphyllium spinithorax Zompro, 2001  [Luzon (Benguet Prov.**)]
Microphyllium haskelli Cumming sp. nov.  [Luzon (Mountain Prov.)]


 • Pseudomicrophyllium Cumming gen. nov.
Pseudomicrophyllium faulkneri Cumming gen. et sp. nov.  [Central Luzon (Ifugao Prov.)]


 • Phyllium Illiger, 1798
Phyllium (Phyllium) ericoriai Hennemann, et al., 2009  [Luzon (Manila, Aurora*, Quirino*, & Quezon Prov.); Batan Is.; Marinduque Is.; Cantanduanes Is.*]
Phyllium (Phyllium) bonifacioi Lit & Eusebio, 2014  [Northern Luzon (Cagayan Prov.* & Northern Luzon***)]
Phyllium (Phyllium) philippinicum Hennemann, et al., 2009****  [Western Luzon (Bataan Prov.)]

Phyllium (Phyllium) antonkozlovi Cumming sp. nov.   [Eastern Luzon (Isabela Prov.)]
Phyllium (Phyllium) bourquei Cumming & Le Tirant sp. nov.  [Luzon (Nueva Vizcaya Prov.)]
Phyllium (Phyllium) geryon Gray, 1843  [Luzon (Aurora Prov.)*]


 Royce T. Cumming, Jing V. Leong and David J. Lohman. 2017. Leaf Insects from Luzon, Philippines, with Descriptions of Four New Species, the New Genus Pseudomicrophyllium, and Redescription of Phyllium (Phyllium) geryon Gray, 1843, (Phasmida: Phylliidae). Zootaxa. 4365(2); 101-131.  DOI: 10.11646/zootaxa.4365.2.1


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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Mollusca • 2017] Like A Bat Out of Heaven: the Phylogeny and Diversity of the Bat-winged Slugs (Heterobranchia: Gastropteridae) ---ScRaBBlE


(B) Gastropteron minutum sp. nov., Maui, Hawai’ian Islands, photo by Cory Pittman. 
(C) Gastropteron multo sp. nov., Mabini, Philippines. 

(E) Sagaminopteron multimaculatum sp. nov., Tingloy, Philippines.
 (F) Siphopteron vermiculum sp. nov., Tingloy, Philippines.

(C) Siphopteron makisig sp. nov., Calatagan, Philippines. 
(D) Siphopteron dumbo sp. nov., Puerto, Galera, Philippines.

Ong & Gosliner, 2017 in Ong, Hallas & Gosliner, 2017. 

Abstract
A molecular phylogeny is presented for 25 newly sequenced specimens of Gastropteridae. The present phylogeny was estimated by analysing the nuclear fragment 28S and two mitochondrial fragments cytochrome c oxidase I (COI) and 16S using maximum likelihood and Bayesian analyses. The distinctness of eight new species of Gastropteridae is supported by the molecular phylogeny and by subsequent Automatic Barcode Gap Discovery (ABGD) analysis. Morphological data also support the distinctness of these species. The following species are described here: Gastropteron minutum Ong and Gosliner sp. nov.Gastropteron multo Ong and Gosliner sp. nov.Sagaminopteron multimaculatum Ong and Gosliner sp. nov., Siphopteron vermiculum Ong and Gosliner sp. nov.Siphopteron flavolineatum Ong and Gosliner sp. nov.Siphopteron nakakatuwa Ong and Gosliner sp. nov.Siphopteron makisig Ong and Gosliner sp. nov. and Siphopteron dumbo Ong and Gosliner sp. nov. All of these species, spanning much of the phylogenetic tree of Gastropteridae, are found in a single, highly diverse region of the Philippines, the Verde Island Passage. These data support the hypothesis that this region is an area of high species richness as well as phyletic diversity. This study also supports strong correlation between morphological characters and the molecular phylogeny within the species of Siphopteron. Molecular studies also indicate the distinctness of specimens of Siphopteron quadrispinosum from Hawaii and those from the western Pacific. Western Pacific specimens should be regarded as Siphopteron leahSiphopteron pohnpei is transferred to Sagaminopteron based on the molecular phylogeny. Other species complexes indicating the presence of geographically separated cryptic species indicate that further detailed study of this group is warranted and that hidden diversity is likely to increase with additional study.


 (A) Gastropteron minutum sp. nov., partaype, CASIZ 199181, Tingloy, Philippines. (B) Gastropteron minutum sp. nov., holotype, CASIZ 209032, Maui, Hawai’ian Islands, photo by Cory Pittman. (C) Gastropteron multo sp. nov., holotype NMP 041181, Mabini, Philippines.

(D) 
Gastropteron multo sp. nov., paratype, CASIZ 186048, Mabini, Philippines. (E) Sagaminopteron multimaculatum sp. nov., holotype, NMP 041182, Tingloy, Philippines. (F) Siphopteron vermiculum sp. nov., Paratype, CASIZ 199129, Tingloy, Philippines.

(A) Siphopteron nakakatuwa sp. nov., partaype, CASIZ 199114, Tingloy, Philippines.  (C) Siphopteron makisig sp. nov., holotype NMP 041186, Calatagan, Philippines. (D) Siphopteron dumbo sp. nov., NMP 041187, Puerto, Galera, Philippines. 

 Photos by T. Gosliner unless otherwise indicated.


Living animals. (A) Gastropteron minutum sp. nov., partaype, CASIZ 199181, Tingloy, Philippines. (B) Gastropteron minutum sp. nov., holotype, CASIZ 209032, Maui, Hawai’ian Islands, photo by Cory Pittman. (C) Gastropteron multo sp. nov., holotype NMP 041181, Mabini, Philippines. (D) Gastropteron multo sp. nov., paratype, CASIZ 186048, Mabini, Philippines. (E) Sagaminopteron multimaculatum sp. nov., holotype, NMP 041182, Tingloy, Philippines. (F) Siphopteron vermiculum sp. nov., Paratype, CASIZ 199129, Tingloy, Philippines. (Photos by T. Gosliner unless otherwise indicated.) 

Living animals. (A) Siphopteron nakakatuwa sp. nov., partaype, CASIZ 199114, Tingloy, Philippines. (B) Siphopteron nakakatuwa sp. nov., holotype, NMP 041185, Tingloy, Philippines. (C) Siphopteron makisig sp. nov., holotype NMP 041186, Calatagan, Philippines. (D) Siphopteron dumbo sp. nov., NMP 041187, Puerto, Galera, Philippines.  (G) Siphopteron flavolineatum, holotype, NMP 041184, Tingloy, Philippines. (H) Siphopteron flavolineatum, paratype, CASIZ 199132, Tingloy, Philippines. Photos by T. Gosliner. 

CONCLUSIONS: 
Based on our morphological studies and molecular phylogeny, we confirmed that the eight new species hypothesized from our field observations of living animals all represent new species that are described here. Our molecular phylogeny also indicates the likely existence of geographically isolated cryptic species within previously classified specimens. Sagaminopteron psychedelicum from Madagascar appears to represent a cryptic species distinct from western Pacific populations, as is Sagaminopteron nigropunctatum from Saudi Arabia. Also, the western Indian Ocean specimen of S. tigrinum represents a cryptic species distinct from the western Pacific S. tigrinum. The lack of strong support for the monophyly of Gastropteron and Siphopteron requires additional study of taxa to determine whether greater support of these lineages is found when taxon sampling is increased. Also the phylogenetic and systematic relationship of Enotepteron needs to be examined when members of this taxon become available for molecular studies.


Elise Ong, Joshua M Hallas and Terrence M Gosliner. 2017. Like A Bat Out of Heaven: the Phylogeny and Diversity of the Bat-winged Slugs (Heterobranchia: Gastropteridae).  Zoological Journal of the Linnean Society. 180(4); 755–789.   DOI: 10.1093/zoolinnean/zlw018 

     

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Herpetology • 2018] Cyclocorinae (New Subfamily) • Discovery of An Old, Archipelago-wide, Endemic Radiation of Philippine Snakes ---ScRaBBlE


Subfamily Cyclocorinae

 Weinell & Brown, 2017.

Highlights
• Three snake genera placed into molecular phylogeny for first time
• Hypothesized close relationship between Myersophis and Oxyrhabdium confirmed
• Hypothesized close relationship between Cyclocorus and Hologerrhum confirmed
• Novel, Philippine-endemic clade of snakes described as new subfamily Cyclocorinae
• Diversification within Cyclocorinae inconsistent with Pleistocene “species-pump”

Abstract
The extraordinarily rich land vertebrate biodiversity of the Philippines includes at least 112 species of terrestrial snakes (74% of which are endemic to the archipelago) in 41 genera (12% endemic). Endemic Philippine snake genera include Cyclocorus (two species), Hemibungarus (three species), Hologerrhum (two species), Oxyrhabdium (two species), and Myersophis (monotypic). Although Hemibungarus and Oxyrhabdium have been included in previous species-level phylogenies, the affinities of the other three Philippine endemic genera are completely unknown. We generated novel DNA sequences for six species from four genera and analyzed these in conjunction with data from earlier studies to infer a phylogeny for the group containing Colubridae, Elapoidea (Elapidae + Lamprophiidae), and Homalopsidae. We present a novel phylogenetic result that strongly supports the existence of an entirely endemic Philippine radiation of elapoid snakes that originated 35–25 million years ago. We provide a revised, phylogeny-based classification to accommodate the new clade, transfer CyclocorusHologerrhum, and Myersophis to Lamprophiidae, and provide the first estimate of the evolutionary relationships among these genera and the related Oxyrhabdium, setting the stage for future investigation of this entirely endemic, novel Philippine elapoid radiation.

Keywords: Cyclocorinae; CyclocorusHologerrhum; Lamprophiidae; MyersophisOxyrhabdium

Representative taxa of the Philippine elapoid radiation, subfamily Cyclocorinae: 
 
Myersophis alpestris (holotype, KU 203012),  Oxyrhabdium modestum (RMB 19937),Cyclocorus lineatus (KU 275744), Oxyrhabdium leporinum leporinum (KU 322339)

[map] Geographic distribution of Cyclocorinae new subfamily in the Philippines, including Cyclocorus, Hologerrhum, Myersophis, Oxyrhabdium, and the unnamed lineage.

Fig. 1. Representative taxa of the Philippine elapoid radiation, subfamily Cyclocorinae:
(A) Oxyrhabdium leporinum leporinum (KU 322339), (B) Oxyrhabdium leporinum visayanum, (C) Oxyrhabdium modestum (Uncataloged KU specimen [RMB 19937]), (D) Hologerrhum philippinum (KU 330056), (E) Cyclocorus lineatus (KU 275744), (F) Cyclocorus nuchalis (KU 344159), (G) and (H) Myersophis alpestris (holotype, KU 203012).

Cyclocorinae (new subfamily)

Type genus: Cyclocorus Duméril, Bibron, and Duméril, 1854
Type species: Lycodon lineatus Reinhardt, 1843

Content: Cyclocorinae includes four endemic Philippine genera:
Genus CyclocorusCyclocorus lineatus lineatusC. l. alcalaiC. nuchalis nuchalis and C. n. taylori.
Genus HologerrhumHologerrhum philippinum and H. dermali.
Genus OxyrhabdiumOxyrhabdium leporinum leporinum, O. l. visayanum and O. modestum.
Genus MyersophisMyersophis alpestris.






Jeffrey L. Weinell and Rafe M. Brown. 2018. Discovery of An Old, Archipelago-wide, Endemic Radiation of Philippine Snakes. Molecular Phylogenetics and Evolution. In Press.   DOI: 10.1016/j.ympev.2017.11.004

  


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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Herpetology • 2015] Platymantis quezoni • A New Species of Limestone Karst inhabiting Forest Frog, Genus Platymantis (Ceratobatrachidae: subgenus Lupacolus) from southern Luzon Island, Philippines ---ScRaBBlE


Platymantis (Lupacolusquezoni 
Brown, de Layola, Lorenzo, Diesmos & Diesmos, 2015 


Abstract

We describe a new species of limestone karst dwelling forest frog of the genus Platymantis from the Quezon Protected Landscape in southeastern Luzon Island, Philippines. We assign Platymantis quezoni, sp. nov., to the diverse assemblage of terrestrial species in the Platymantis dorsalis Group, subgenus Lupacolus on the basis of its body size and proportions, only slightly expanded terminal discs of the fingers and toes, and its terrestrial microhabitat. The new species is distinguished from these and all other Philippine congeners by features of its external morphology, its restriction to a distinctive limestone karst microhabitat, and its advertisement call, which is unique among frogs of the family Ceratobatrachidae. Several distinguishing morphological characters include its moderate body size (22.1–33.9 mm SVL for 16 adult males and 32.4–39.7 mm SVL for five adult females), slightly expanded terminal discs of the fingers and toes, smooth skin with limited dermal tuberculation, and a dorsal color pattern of mottled tan to dark brown with black blotches. The new species is the sixth Philippine Platymantis known to occur exclusively on limestone karst substrates (previously known karst-obligate species include: P. bayani, P. biak, P. insulatus, P. paengi, and P. speleaus). Recently accelerated discovery of limestone karst anurans across the Philippines suggests that numerous additional species may await discovery on the hundreds of scattered karst formations throughout the archipelago. This possibility suggests that a major conservation priority in coming years will be to study, characterize, describe, and preserve the endemic species supported by this patchy, unique and imperiled type of forest ecosystem in the Philippines.

Keywords: Amphibia, Biodiversity; Evolutionary convergence; Limestone ecomorph; Microhabitat preference; Species discovery


FIGURE 3. Photographs of Platymantis (Lupacolus) quezoni, sp. nov.,
(A) the holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3).
Photos: R. M. Brown


FIGURE 5. Limestone karst forest habitat of Platymantis (Lupacolusquezonisp. nov., at the type locality:
 (A) limestone bolder where the holotype (PNM 9817) was recorded and collected; (B) appearance of habitat in late afternoon (~16:30). Note darkened caves and crevices beneath limestone over-hangs from where P. quezoni initiate vocalizations in the late afternoons (16:30–17:30 hr) from concealed perches inside crevices and small caves.
 Photos: R. M. Brown. 


Distribution. Platymantis quezoni is known only from the type locality but may be more wide spread in scattered karst outcrops of northern portions of the Bicol Peninsula (Restificar et al. 2006; MGB 2010), especially on limestone karst formations in the nearby Bondoc Peninsula (Fig. 1). 

 Etymology.  The specific epithet is a patronym in the genitive singular, honoring Manuel Luis Molina Quezon. Quezon served as president of the Commonwealth of the Philippines during the American colonial period from 1935 through the conclusion of the Second World War. An exemplary statesman, he led the struggle for Philippine independence from American rule.
Suggested common name: Quezon Limestone Forest Frog.  



 Rafe M. Brown, Louise Abigail Asio De Layola, Antonio Ii Nuñez Lorenzo, Mae Leonida Diesmos and Arvin Cantor Diesmos. 2015. A New Species of Limestone Karst inhabiting Forest Frog, Genus Platymantis (Amphibia: Anura: Ceratobatrachidae: subgenus Lupacolus) from southern Luzon Island, Philippines. Zootaxa. 4048(2): 191–210.  DOI: 10.11646/zootaxa.4048.2.3



    


---------------------------------------------------------------
روابط التحميل والمشاهدة، الروابط المباشرة للتحميل
او
شاهد هذا الفيديو القصير لطريقة التحميل البسيطة


كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

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