[PaleoMammalogy • 2017] Mystacodon selenensis • Earliest Mysticete from the Late Eocene of Peru Sheds New Light on the Origin of Baleen Whales ---ScRaBBlE

Mystacodon selenensis 

Lambert, Martínez-Cáceres, Bianucci, Celma, Salas-Gismondi, Steurbaut, Urbina & Muizon, 2017

 DOI:  10.1016/j.cub.2017.04.026 

Highlights

• An ancient whale is described based on a skeleton from the late Eocene of Peru

• It is identified as the earliest known mysticete (baleen whales and relatives)

• Skeletal anatomy provides crucial information on archaeocete-mysticete transition

• This whale is interpreted as specialized for suction and possibly benthic feeding

Summary

Although combined molecular and morphological analyses point to a late middle Eocene (38–39 million years ago) origin for the clade Neoceti (Odontoceti, echolocating toothed whales plus Mysticeti, baleen whales, and relatives), the oldest known mysticete fossil dates from the latest Eocene (about 34 million years ago) of Antarctica. Considering that the latter is not the most stemward mysticete in recent phylogenies and that Oligocene toothed mysticetes display a broad morphological disparity most likely corresponding to contrasted ecological niches, the origin of mysticetes from a basilosaurid ancestor and its drivers are currently poorly understood. Based on an articulated cetacean skeleton from the early late Eocene (Priabonian, around 36.4 million years ago) of the Pisco Basin, Peru, we describe a new archaic tooth-bearing mysticete, Mystacodon selenensis gen. et sp. nov. Being the geologically oldest neocete (crown group cetacean) and the earliest mysticete to branch off described so far, the new taxon is interpreted as morphologically intermediate between basilosaurids and later toothed mysticetes, providing thus crucial information about the anatomy of the skull, forelimb, and innominate at these critical initial stages of mysticete evolution. Major changes in the morphology of the oral apparatus (including tooth wear) and flipper compared to basilosaurids suggest that suction and possibly benthic feeding represented key, early ecological traits accompanying the emergence of modern filter-feeding baleen whales’ ancestors.

Systematics

Cetacea

Pelagiceti

Neoceti

Mysticeti

Mystacodontidae fam. nov.

Mystacodon selenensis gen. et sp. nov.

Etymology: From ancient Greek mystacos (“moustache”) in reference to the suborder Mysticeti and odontos (“tooth”), “mysticete with teeth,” and from Selene, the Greek goddess of the moon, in reference to the Playa Media Luna type locality.

Figure 1: Cranium, Mandible, and Teeth of Mystacodon selenensis gen. et sp. nov. MUSM 1917.

Cranium in dorsal (A), left lateral (B), and ventral (C) views; detail of left posterior lower teeth in lateral view (D); left mandible in lateral (E) and dorsal (F) views; and three detached anterior lower teeth (from left to right: incisor, incisor/canine, and ?p1) in lingual or labial and occlusal views (G).

aof, antorbital foramina; apm, antorbital process of maxilla; bn, bony nares; C, upper canine; c, lower canine; cp, coronoid process; ep, embrasure pit; I1–I3, upper incisors; i1–i3, lower incisors; iop, infraorbital plate; ju, jugal; la, lacrimal; M1 and M2, upper molars; m1–m3, lower molars; maf, mandibular fossa; mf, mental foramina; mg, mesorostral groove; mx, maxilla; na, nasal; nc, nuchal crest; P1–P4, upper premolars; p1–p4, lower premolars; pa, parietal; pmx, premaxilla; prpf, preorbital process of frontal; pspf, postorbital process of frontal; sq, squamosal; sym, mandibular symphysis; zpm, zygomatic process of maxilla; zyg, zygomatic process of squamosal. Scale bars for (A)–(C), (E), and (F), 200 mm; for (D), 20 mm; and for (G), 10 mm.   

Two Mystacodon selenensis individuals diving down to catch eagle rays along the seafloor of a shallow cove off the coast of present-day Peru.

Illustration: Alberto Gennari 

Holotype: Museo de Historia Natural, Universidad Nacional Mayor de San Marcos (MUSM; Lima, Peru) 1917, partial skeleton including cranium, mandibles, teeth, cervical, thoracic, lumbar and caudal vertebrae, ribs, partial right and left forelimbs, and left innominate.

Locality: Playa Media Luna, southern part of Pisco Basin, southern coast of Peru, ...

Horizon: Middle part of the Yumaque Formation, 77 m above the base; lower part of calcareous nannofossil zone NP19/20 of Martini; dated at 36.4 million years ago based on age estimations used by Agnini et al. [2014]; early late Eocene (early Priabonian; see Figures S1A and S1D, Table S1, and STAR Methods for the biostratigraphic and biochronological interpretations).

Diagnosis: MUSM 1917 is identified as a Neoceti based on the following derived characters, absent in basilosaurid archaeocetes: partly open mesorostral groove; anteroposteriorly elongated rostral portion of maxilla; loss of sagittal crest; supraoccipital shield anterodorsally inclined; apex of zygomatic process of squamosal nearly contacting postorbital process of frontal; and distal epiphysis of the humerus divided in two angled radial and ulnar facets. It can be referred to the Mysticeti due to the following combination of derived characters: dorsoventrally thin lateral edge of maxilla on rostrum; presence of an antorbital process of the maxilla; presence of a maxillary infraorbital plate; and triangular supraoccipital shield. It is further diagnosed by two possibly autapomorphic features: nasal anteroposteriorly longer than frontal plus parietal and strong tuberosity on anterior edge of radius; two additional derived characters: posteriormost upper tooth anterior to level of antorbital process of maxilla and broad-based rostrum (ratio between width of skull at rostrum base and width at postorbital process > 0.8); and a series of plesiomophic features: supraoccipital shield not extending anterior to anterior level of squamosal fossa, only two dorsal infraorbital foramina, a basilosaurid dental formula 3.1.4.2/3.1.4.3, no wide diastemata between posterior cheek teeth, sutured mandibular symphysis, and well-defined acetabulum on innominate. Finally, MUSM 1917 lacks cranial synapomorphies of Odontoceti: facial concavity, presence of premaxillary foramen and premaxillary sac fossa, and posterior expansion of maxilla over the supraorbital region (Figures 1, 2, 3, and S2).

Members of the excavation team digging around the skeleton of Mystacodon selenensis at the Media Luna locality in the Pisco Basin, Peru.

photo: Giovanni Bianucci  

Olivier Lambert, Manuel Martínez-Cáceres, Giovanni Bianucci, Claudio Di Celma, Rodolfo Salas-Gismondi, Etienne Steurbaut, Mario Urbina and Christian de Muizon. 2017. Earliest Mysticete from the Late Eocene of Peru Sheds New Light on the Origin of Baleen Whales. Current Biology.  In Press.  DOI:  10.1016/j.cub.2017.04.026

    

36m-year-old fossil discovery is missing link in whale evolution, say researchers TheGuardian.com/science/2017/may/11/36m-year-old-fossil-discovery-is-missing-link-in-whale-evolution-say-researchers-mystacodon-selenensis

Baleen whales' ancestors were toothy suction feeders http://phy.so/413697728 via @physorg_com

This ancient whale likely sucked prey into its mouth like a giant vacuum cleaner https://www.theverge.com/tldr/2017/5/11/15610764/mysticete-whale-ancestor-suction-feeder-evolution  via @Verge

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Herpetology • 2018] Cyclocorinae (New Subfamily) • Discovery of An Old, Archipelago-wide, Endemic Radiation of Philippine Snakes ---ScRaBBlE

Subfamily Cyclocorinae

 Weinell & Brown, 2017.

   DOI: 10.1016/j.ympev.2017.11.004 

Highlights

• Three snake genera placed into molecular phylogeny for first time

• Hypothesized close relationship between Myersophis and Oxyrhabdium confirmed

• Hypothesized close relationship between Cyclocorus and Hologerrhum confirmed

• Novel, Philippine-endemic clade of snakes described as new subfamily Cyclocorinae

• Diversification within Cyclocorinae inconsistent with Pleistocene “species-pump”

Abstract

The extraordinarily rich land vertebrate biodiversity of the Philippines includes at least 112 species of terrestrial snakes (74% of which are endemic to the archipelago) in 41 genera (12% endemic). Endemic Philippine snake genera include Cyclocorus (two species), Hemibungarus (three species), Hologerrhum (two species), Oxyrhabdium (two species), and Myersophis (monotypic). Although Hemibungarus and Oxyrhabdium have been included in previous species-level phylogenies, the affinities of the other three Philippine endemic genera are completely unknown. We generated novel DNA sequences for six species from four genera and analyzed these in conjunction with data from earlier studies to infer a phylogeny for the group containing Colubridae, Elapoidea (Elapidae + Lamprophiidae), and Homalopsidae. We present a novel phylogenetic result that strongly supports the existence of an entirely endemic Philippine radiation of elapoid snakes that originated 35–25 million years ago. We provide a revised, phylogeny-based classification to accommodate the new clade, transfer Cyclocorus, Hologerrhum, and Myersophis to Lamprophiidae, and provide the first estimate of the evolutionary relationships among these genera and the related Oxyrhabdium, setting the stage for future investigation of this entirely endemic, novel Philippine elapoid radiation.

Keywords: Cyclocorinae; Cyclocorus; Hologerrhum; Lamprophiidae; Myersophis; Oxyrhabdium

Representative taxa of the Philippine elapoid radiation, subfamily Cyclocorinae: 
 Myersophis alpestris (holotype, KU 203012),  Oxyrhabdium modestum (RMB 19937),Cyclocorus lineatus (KU 275744), Oxyrhabdium leporinum leporinum (KU 322339)

[map] Geographic distribution of Cyclocorinae new subfamily in the Philippines, including Cyclocorus, Hologerrhum, Myersophis, Oxyrhabdium, and the unnamed lineage.

  DOI: 10.1016/j.ympev.2017.11.004  

Fig. 1. Representative taxa of the Philippine elapoid radiation, subfamily Cyclocorinae:
(A) Oxyrhabdium leporinum leporinum (KU 322339), (B) Oxyrhabdium leporinum visayanum, (C) Oxyrhabdium modestum (Uncataloged KU specimen [RMB 19937]), (D) Hologerrhum philippinum (KU 330056), (E) Cyclocorus lineatus (KU 275744), (F) Cyclocorus nuchalis (KU 344159), (G) and (H) Myersophis alpestris (holotype, KU 203012).

Cyclocorinae (new subfamily)

Type genus: Cyclocorus Duméril, Bibron, and Duméril, 1854

Type species: Lycodon lineatus Reinhardt, 1843

Content: Cyclocorinae includes four endemic Philippine genera:

Genus Cyclocorus: Cyclocorus lineatus lineatus, C. l. alcalai, C. nuchalis nuchalis and C. n. taylori.

Genus Hologerrhum: Hologerrhum philippinum and H. dermali.

Genus Oxyrhabdium: Oxyrhabdium leporinum leporinum, O. l. visayanum and O. modestum.

Genus Myersophis: Myersophis alpestris.

Jeffrey L. Weinell and Rafe M. Brown. 2018. Discovery of An Old, Archipelago-wide, Endemic Radiation of Philippine Snakes. Molecular Phylogenetics and Evolution. In Press.   DOI: 10.1016/j.ympev.2017.11.004

  

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Daliansaurus liaoningensis • A New Troodontid Dinosaur from the Lower Cretaceous Yixian Formation of Liaoning Province, China ---ScRaBBlE

Daliansaurus liaoningensis 

Shen, Lü, Liu, Kundrát, Brusatte & Gao, 2017

GeoJournals.cn/dzxben

Abstract

 A new troodontid dinosaur, Daliansaurus liaoningensis gen. et sp. nov., is erected based on a nearly complete specimen from the Lower Cretaceous Yixian Formation of Beipiao City, Liaoning Province, China. This well preserved skeleton provides important new details of the anatomy for Liaoning troodontids, and gives new insight into their phylogenetic relationships and evolution. Daliansaurus is distinguished from other troodontids by an enlarged ungual on pedal digit IV, which is approximately the same size as the sickle-shaped second ungual, and is differentiated from other Liaoning troodontids by a number of characters of the skull, manus, pelvis, and hindlimb. A phylogenetic analysis recovers Daliansaurus within a subclade of Liaoning troodontids that also includes Sinovenator, Sinusonasus, and Mei. We erect a name for this group — Sinovenatorinae — and argue that it reflects a localized radiation of small-bodied troodontids in the Early Cretaceous of eastern Asia, similar to previously recognized radiations of Liaoning dromaeosaurids and avialans. As more Liaoning theropods are discovered, it is becoming apparent that small, feathered paravians were particularly diverse during the Early Cretaceous, and future work is needed to clarify how this diversity arose, which species coexisted, and how these numerous species partitioned niches. 

Key words: Troodontids, Daliansaurus liaoningensis, Early Cretaceous, Yixian Formation, Liaoning

Fig. 15. The living scene of Daliansaurus liaoningensis gen. et sp. nov. (DNHM D 2885) (drawn by Zhao Chuang)
Fig. 2. Photograph of the holotype of Daliansaurus liaoningensis gen. et sp. nov. (DNHM D2885).

Theropoda Marsh, 1881

Maniraptora Gauthier, 1986

Troodontidae Gilmore, 1924 

 Sinovenatorinae subfamily nov. 

Definition: The most inclusive clade including Sinovenator changii but not Troodon formosus, Saurornithoides mongoliensis, Anchiornis huxleyi, Archaeopteryx lithographica, Gallus gallus, Unenlagia comahuensis, or Dromaeosaurus albertensis. 

  Diagnosis: Based on our phylogenetic analysis, this clade can be diagnosed with the following combination of characters (for further discussion, see below): a premaxilla whose anteroposterior length is less than 10% of the length of the maxilla along the ventral margin of the upper jaw; a premaxillary maxillary process that contacts the nasal ventral to the external naris; some maxillary and dentary teeth that lack serrations on the mesial carina; unfused zygapophyses on the sacral vertebrae; an antitrochanter that is posterodorsal to the acetabulum; and a metatarsal IV that is mediolaterally wide and flattened in cross section.

 Included species: Based on our phylogenetic analysis, this clade currently includes Sinovenator changii, Sinusonasus magnodens, Mei long, and Daliansaurus liaoningensis gen. et sp. nov.

 Daliansaurus liaoningensis gen. et sp. nov.

Etymology: The generic name refers to the city of Dalian in the south of Liaoning province. The specific name “liaoningensis” refers to the province in which the specimen was discovered.

Locality and horizon: Lujiatun, Beipiao City, Liaoning Province; Yixian Formation, Lower Cretaceous.

Fig. 15. The living scene of Daliansaurus liaoningensis gen. et sp. nov. (DNHM D 2885) (drawn by Zhao Chuang) 

Conclusions

 It appears that small-bodied, feathered, winged paravian theropods from Liaoning—dromaeosaurids, avialans, and troodontids—were exceptionally diverse. Each group experienced its own local Early Cretaceous diversification, which produced a number of species that either coexisted or, perhaps more likely, replaced each other over time or partitioned niches and geographic ranges so that they did not extensively overlap in space. The discovery of Daliansaurus also demonstrates that Liaoning continues to produce new dinosaur taxa, and remains one of the world’s most productive sites for understanding paravian evolution, more than two decades after the first feathered dinosaurs were reported.

 SHEN Caizhi, LÜ Junchang, LIU Sizhao, Martin KUNDRÁT, Stephen L. BRUSATTE and GAO Hailong. 2017. A New Troodontid Dinosaur from the Lower Cretaceous Yixian Formation of Liaoning Province, China.  ACTA GEOLOGICA SINICA.  91(3); 763–780. 

http://www.geojournals.cn/dzxben/ch/reader/view_abstract.aspx?file_no=2017endzxb03001&flag=1

http://www.geojournals.cn/dzxben/ch/reader/create_pdf.aspx?file_no=2017endzxb03001&flag=1&journal_id=dzxben&year_id=2017

twitter.com/SteveBrusatte/status/873526560564944898

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Ichthyology • 2017] Tarumania walkerae • Tarumaniidae fam. nov., A New Family of Neotropical Freshwater Fishes from Deep Fossorial Amazonian Habitat, with A Reappraisal of Morphological Characiform Phylogeny (Teleostei: Ostariophysi) ---ScRaBBlE

Tarumania walkerae

de Pinna, Zuanon, Py-Daniel & Petry, 2017

 DOI: 10.1093/zoolinnean/zlx028  

Abstract

We present the taxonomic description of Tarumaniidae, a new family of South American freshwater fishes including a new genus (Tarumania) and species (Tarumania walkerae) from the Central Amazon. The new taxon displays an extraordinary set of unique characteristics, which sets it apart from all other known bony fishes, either in South America or elsewhere. Tarumaniidae has reverse-imbricated scales on the head, 244 or more scales along the midlateral lateral series, an 11-chambered swimbladder extending along most of the body, an anteriorly deflectable pelvic fin and a platybasic skull, among other unusual traits. Although the general habitus of Tarumania gives little hint of its phylogenetic placement, examination of internal anatomy is largely elucidative concerning the main features of its relationships. For example, Tarumania has a complete Weberian apparatus, comprising a set of characteristics supporting its position in Otophysi and Ostariophysi. Additional comparative evidence corroborates its placement in the order Characiformes, including some trademark characiform synapomorphies such as the presence of a hypertrophied lagenar capsule. We conducted a phylogenetic analysis on the basis of previously published morphological evidence for Characiformes, with the inclusion of Tarumania and additional relevant taxa and characters. Results support the position of Tarumania as a part of the superfamily Erythrinoidea and sister group to the neotropical Erythrinidae. Our increased taxonomic representation of Erythrinidae in the matrices firmly corroborates Tarumania as sister group to the whole of Erythrinidae and not to a subgroup of that family. Tarumania walkerae is a predator, which inhabits vast leaf-litter deposits in the Rio Negro drainage, and is found deeply buried in isolated pools during the dry season. It is the first characiform with an anguilliform body shape and fossorial habits. The unprecedented combination of phenotypic traits, behaviour and habitat adds substantially to the larger picture of freshwater fish diversity. Although its microhabitat is difficult to sample, the geographical range of T. walkerae includes areas close to major urban centres and which have been ichthyologically surveyed for many years. The fact that a relatively large and highly distinctive fish such as Tarumania remained undiscovered until now is testimony to the still-incomplete state of knowledge of biodiversity in Amazonian waters.

Keywords: Amazon, Characiformes, Erythrinoidea, new genus, new species, phylogeny, systematics, taxonomy

Figure 1. Tarumania walkerae gen. et sp. nov., holotype, INPA 33737. 

Figure 2. Tarumania walkerae gen. et sp. nov., holotype, INPA 33737. 
Figure 3. Live specimen, juvenile, of Tarumania walkerae, paratype, MZUSP 120543, shortly after collection. Arrow shows pelvic fins in anteriorly deflected position. 

TAXONOMIC ACCOUNTS

Tarumaniidae fam. nov.

Tarumania gen. nov. 
Type species: Tarumania walkerae sp. nov. 

Etymology: From the river Tarumã-Mirim, tributary of the lower Rio Negro, first known locality of the new taxon. A noun in nominative singular. Gender feminine.

Tarumania walkerae sp. nov.

Etymology: The specific name honours eminent limnologist Ilse Walker [Instituto Nacional de Pesquisas da Amazônia, Manaus (INPA)], not only for her lifelong contribution to the knowledge of Amazonian ecology but also for having collected the first (and for some years, only) known specimen. The epithet is a noun in the genitive feminine case.

Geographical distribution: So far endemic to the Rio Negro basin, from the Rio Tarumã-Mirim, tributary to the Rio Negro near the city of Manaus, and from the Anavilhanas archipelago, on the main Rio Negro (Fig. 11). The two localities are ~60 km apart in straight line.

Ecological notes All specimens known of Tarumania walkerae were collected deeply buried into massive leaf-litter deposits in isolated pools in the riparian forest. ....

Figure 2. Tarumania walkerae gen. et sp. nov., holotype, INPA 33737. Dorsal (a) and ventral (b) views of head.
Figure 5. Tarumania walkerae, paratype, INPA 21603, lateral view of head showing reverse-imbricated scales. Specimen cleaned of superficial mucus.

Mário de Pinna, Jansen Zuanon, Lucia Rapp Py-Daniel and Paulo Petry. 2017. A New Family of Neotropical Freshwater Fishes from Deep Fossorial Amazonian Habitat, with A Reappraisal of Morphological Characiform Phylogeny (Teleostei: Ostariophysi). Zoological Journal of the Linnean Society. DOI:   10.1093/zoolinnean/zlx028 

facebook.com/SouthAmericanIchthyology/photos/1620698521284272
Paulo and the Mysterious Muck Fish youtu.be/FIzUGi7k9M0 

  

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[PaleoIchthyology • 2016] Eekaulostomus cuevasae • An Ancient Armored Trumpetfish (Aulostomoidea) from Danian (Paleocene) Marine Deposits of Belisario Domínguez, Chiapas, southeastern Mexico ---ScRaBBlE

Eekaulostomus cuevasae 

Cantalice & Alvarado-Ortega,  2016

 palaeo-electronica.org

ABSTRACT

Eekaulostomus cuevasae gen. and sp. nov. is described and identified here as a new member of the superfamily Aulostomoidea. The single specimen known of this species is part of a newly fossil assemblage collected in the marine sediments belonging to the early Paleocene Tenejapa-Lacandón geological unit, exploited in the Belisario Domínguez quarry, near Palenque town, State of Chiapas, southeastern Mexico. E. cuevasae represents the oldest aulostomoid as far known and the first fossil species of this superfamily collected in America. E. cuevasae differs from other aulostomoids in the presence of two spines preceding the soft rays of both dorsal and anal fins; the star-like scales covering the entire body and part of the snout; as well as the relative large number of principal rays in the caudal fin. The recognition of E. cuevasae as the stem group of Aulostomoidea increases the temporal and geographic distribution of this superfamily up to Danian and within the Caribbean region, when a large part of Chiapas was under the sea after the Cretaceous-Tertiary mass extinction event. This finding also provides evidences suggesting the membership of Aulostomoidea within the order Gasterosteiformes, in which the scutes covering the trunk and the robust spines in unpaired fins are recurrent features.

  Keywords: new species; Aulostomoidea; fossil; Paleocene; Chiapas; Mexico

Figure 2: Holotype. IGM 4716, almost complete specimen exposing the right lateral side of the body.  

Systematic Paleontology

Superfamily AULOSTOMOIDEA sensu Greenwood et al., 1966

Family EEKAULOSTOMIDAE fam. nov.

Genus EEKAULOSTOMUS gen. nov.

Type species. Eekaulostomus cuevasae sp. nov.,  

Derivation of name. The genus name includes the Mayan word “Eek” (= star), the Greek word "aulos" (= αὐλός, that is the name of an ancient flute), and the Latin word "stoma" (= mouth). The name refers to a "fish with a star-like scutes and flute-shaped mouth."

Eekaulostomus cuevasae sp. nov.

Derivation of name. The specific epithet of this fish honors our colleague, Martha Cuevas García, whose dedication and newly passion for the fossils led us to find the only specimen of Eekaulostomus cuevasae.

Occurrence. Paleocene (Danian, ≈ 63Ma) marine strata of the Tenejapa-Lacandón geological unit. Belisario Domínguez quarry, Salto de Agua Municipality, State of Chiapas, southeastern Mexico (Alvarado-Ortega et al., 2015).

Diagnosis. Aulostomoidea fish with rigid star-like scutes covering the whole trunk and part of the snout; pelvic fin placed anteriorly, just behind the postcleithrum; two spines in front of the soft rays of dorsal and anal fins; eight soft rays in both anal and dorsal fins; caudal fin formula iv+I+7—5+I+iii.

CONCLUSIONS 

Fossils referring to the superfamily Aulostomoidea had been collected more than 200 years ago in Eocene and younger marine deposits along Europe. Although the extant aulostomoids form part of large modern cladistics essays, some are based on morphological evidences and others on molecular data; unfortunately, the fossil aulostomoids have never been phylogenetically studied. This situation has prevented the generation of a robust classification of the aulostomoids, and at the same time, has fueled the differences and contradictions between the phylogenetic hypotheses already published. It is so, that it is desirable to make these European fossils part of future cladistic studies; however, first we have to fulfill the task of re-describing them accurately using modern and homogeneous criteria. Only up to the present day, the distribution of fossil aulostomoids was restricted to Europe. Although this fact has not interested paleontologists outside of Europe to further collaborate in studies concerning the diversity evolution of aulostomoids; the goal of this paper is to provide the first tangible evidence that in the past, this fish group was also an inhabitant of the American seas.

From now on, we must take more seriously the paleontological surveys on late Cretaceous and early Paleocene sites with marine sediments present throughout the tropical region of America. As present study shows, fossils may exist that allow us to delve into the details of the evolution of the fishes on both sides of Cretaceous-Tertiary mass extinction event. Sadly, during the joint INAH-UNAM project, from which this article was drawn, no other fossil aulostomoid was recovered; however, the collection effort applied in Belisario Domínguez (where Eekaulostomus cuevasae gen. and sp. nov. came) as well as in its coeval and neighbor quarry, División del Norte, both with Danian marine sediments, really is far from reaching saturation.

The re-examination of the relationships of the Aulostomoidea executed here, using datasets previously generated by other authors and including Eekaulostomus cuevasae gen. and sp. nov., might not be the best way to achieve the desired understanding on the evolutionary processes of these fishes; however, this exercise significantly contributes to this goal. On the one side, the undeniable position of this new species as a primitive aulostomoid member, together with its deep morphological differences with other extinct and living taxa formally, or putatively, included in such superfamily, trace possible trends in morphological changes experienced by these fishes since the Paleocene to the present. This essay also widens the geographical scenario where the evolution of these fishes took place, extending from Europe to the tropical region of America.

  Cantalice, Kleyton Magno and Alvarado-Ortega, Jesús. 2016. Eekaulostomus cuevasae gen. and sp. nov., An Ancient Armored Trumpetfish (Aulostomoidea) from Danian (Paleocene) Marine Deposits of Belisario Domínguez, Chiapas, southeastern Mexico. Palaeontologia Electronica. 19.3.53A: 1-24

palaeo-electronica.org/content/2016/1659-ancient-armored-trumpetfish

blogs.plos.org/paleocomm/2016/12/16/fossil-friday-roundup-december-16-2016/

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[PaleoEntomology • 2017] Mesosticta davidattenboroughi • Mesostictinae subfam. nov., An Archaic Group of Platystictid Damselflies (Odonata: Zygoptera) from mid-Cretaceous Burmese Amber ---ScRaBBlE

Mesosticta davidattenboroughi

 Zheng, Wang, Nel, Jarzembowski, Zhang & Chang, 2017

 DOI: 10.1080/14772019.2017.1348395 

Abstract

Odonatans are quite rare in the fossil record compared with the other insects, especially in Cretaceous amber inclusions. The extant family Platystictidae is one of the most diverse Zygoptera, but short of fossil records. In this paper, a new species, Mesosticta davidattenboroughi sp. nov., is described from mid-Cretaceous Burmese amber, representing the third-known fossil species of Platystictidae. Mesosticta davidattenboroughi sp. nov. has a long IR1 beginning one cell distal of the base of RP2, confirming the previous attribution of Mesosticta Huang, Azar, Cai & Nel, 2015 to Platystictidae. It differs from other species of Mesosticta in having a long IR1 and a basally crossed subdiscoidal cell. The fossil genus Mesosticta shares the diagnostic characters of the modern platystictid genera, viz. a basally recessed ‘CuP’ (shared by all species), a very long IR1 (only in Mesosticta davidattenboroughi sp. nov.), and a specialized subdiscoidal area mostly rhomboidal in shape (only in Mesosticta electronica Zheng, Zhang, Chang & Wang, 2016). Based on the platystictid damselflies from Burmese amber, a new subfamily Mesostictinae subfam. nov. is established. Mesostictinae subfam. nov. represents the first fossil group of modern platystictid damselflies, documenting the appearance of Platystictidae as early as mid-Cretaceous. It differs from modern Platystictidae by the presence of fewer postnodal and postsubnodal crossveins, a short MP, the base of RP2 being nearer to the subnodus and the nodus lying more distally.

Keywords: Platystictidae, Zygoptera, Odonata, Cenomanian, Cretaceous, Burmese amber

Figure 1. Mesosticta davidattenboroughi sp. nov., holotype, NIGP164541, photograph of specimen. 

Order Odonata Fabricius, 1793

Suborder Zygoptera Selys-Longchamps, 1854

Superfamily Platystictoidea Kennedy, 1920

Family Platystictidae Kennedy, 1920

Subfamily Mesostictinae subfam. nov.

Type genus. Mesosticta Huang, Azar, Cai & Nel, 2015.

Mesosticta Huang, Azar, Cai & Nel, 2015 

Type species. Mesosticta burmatica Huang, Azar, Cai & Nel, 2015. 

Included species. Mesosticta electronica Zheng, Zhang, Chang & Wang, 2016; Mesosticta davidattenboroughi sp. nov. 

Mesosticta davidattenboroughi sp. nov. 

Type species. NIGP164541, two complete forewings attached to body.

 Diagnosis. Forewing characters: IR1 long, originating one cell distal of base of RP2, nearer to N than to Pt; Arc aligned with Ax2; subdiscoidal cell basally crossed by one vein. 

Etymology. In honour of Sir David Attenborough, on his 90th birthday, for his appreciation of dragonflies. 

Locality and horizon. Hukawng Valley, Kachin Province, Myanmar; lowermost Cenomanian, lowermost Upper Cretaceous.

Figure 7. Hypothetical position of Mesosticta davidattenboroughi sp. nov. in phylogenetic tree of Zygoptera. All line drawings are based on forewings (phylogeny based on Dijkstra et al. 2014; line drawing of Sinosticta ogatai Matsuki & Saito, 1996 after Wilson 1997; line drawing of Palaemnema picicaudata Kennedy, 1938 after Kennedy 1938; line drawings of Platysticta deccanensis Laidlaw, 1915 and Protosticta himalaiaca Laidlaw, 1917 after Fraser 1933).

Daran Zheng, Bo Wang, André Nel, Edmund A. Jarzembowski, Haichun Zhang & Su-Chin Chang. 2017. Mesostictinae subfam. nov., An Archaic Group of Platystictid Damselflies (Odonata: Zygoptera) from mid-Cretaceous Burmese Amber.  Journal of Systematic Palaeontology.   DOI: 10.1080/14772019.2017.1348395

David Attenborough gains new species namesake  phy.so/422069741 @physorg_com

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.