The Anthaxia (Anthaxia) midas Kiesenwetter, 1857 species-group is defined and revised. A new species from Turkey, Anthaxia (A.) cebecii sp. nov., is described and compared to its most similar species. A. midas oberthuri Schaefer, 1938 is elevated to the rank of species, and a lectotype is designated. A. (A.) spathuligera Obenberger, 1924 and A. (A.) midas muelleri Obenberger, 1925 are reconfirmed synonyms of A. midas Kiesenwetter, 1857.
All species of the new species-group are illustrated, including the hitherto unknown male of A. (A.) patsyae Baiocchi, 2008, all type specimens and original data labels. In addition to diagnostic characters, informations on the distribution, biology and taxonomic position of each species are also presented, together with a short definition of the new species-group, and a key to its species.
Keywords: Coleoptera, Buprestidae, Anthaxiini, Anthaxia (A.) midas species-group revision, new species, lectotype designation, distribution, bionomy, taxonomy, Palaearctic region
Daniele Baiocchi and Gianluca Magnani. 2018. A Revision of the Anthaxia (Anthaxia) midas Kiesenwetter, 1857 Species-group (Coleoptera: Buprestidae: Anthaxiini). Zootaxa. 4370(3); 201–254. DOI: 10.11646/zootaxa.4370.3.1
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شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
• A molecular systematic study was conducted for the wide-ranging Trachylepis varia complex.
• Phylogenetic analyses support the existence of at least eight species within the Trachylepis varia complex.
• The Southern African members of the Trachylepis varia complex are phenotypically distinct.
• We update the taxonomy for the southern Africa members of the Trachylepis varia complex.
• Diversification within the Trachylepis varia complex began during the mid to late Miocene or early Pliocene.
Abstract
A systematic study of the Trachylepis varia complex was conducted using mitochondrial and nuclear DNA markers for individuals sampled across the species range. The taxonomic history of T. varia has been complicated and its broad geographic distribution and considerable phenotypic variation has made taxonomic revision difficult, leading earlier taxonomists to suggest that T. varia is a species complex. We used maximum likelihood and Bayesian inference to estimate gene trees and a multilocus time-tree, respectively, and we used these trees to identify the major clades (putative species) within T. varia. Additionally, we used morphological and color pattern data to distinguish and revise the taxonomy of the southern African clades. The major clades recovered in the multilocus time-tree were recovered in each of gene trees, although the relationships among these major clades differed across gene trees. Genetic data support the existence of at least eight species within the T. varia complex, each of which originated during the mid to late Miocene or early Pliocene. We focus our systematic discussion on the southern African members of the T. varia complex, revive Trachylepis damarana(Peters, 1870) and T. laevigata(Peters, 1869), and designate lectotypes for T. damarana and T. varia.
We find strong evidence that Trachylepis varia, T. damarana, and T. laevigata are distinct species that occur in southern Africa and that five additional, species-level clades occur north of the Zambezi and Kunene rivers, although future studies are needed to determine whether Trachylepis nyikae and Trachylepis isellii should also be recognized. The allopatric distribution and morphological distinctiveness of T. isellii (Largen and Spawls, 2010) suggests that this species is probably valid and the presence of multiple endemic species on the Nyika Plateau (Poynton, 1997; Burrows and Willis, 2005) suggests that T. nyikae may also be a valid species. Additionally, little is known about the distribution or natural history of the undescribed species sampled in Ethiopia, Democratic Republic of the Congo, or Tanzania. Lastly, next generation DNA sequencing may be useful in resolving deeper phylogenetic relationships within the T. varia complex and for distinguishing historical gene flow from incomplete lineage sorting. This study is the first to use genetic data to address species diversity, phylogenetic history, and taxonomic issues for the T. varia complex and is an example of how both genetic and phenotypic data can be used to resolve taxonomic problems and to estimate species ranges.
Jeffrey L. Weinell and Aaron M. Bauer. 2018. Systematics and Phylogeography of the Widely Distributed African Skink Trachylepis varia Species Complex. Molecular Phylogenetics and Evolution. 120; 103-117. DOI: 10.1016/j.ympev.2017.11.014
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شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
Specimens with intermediate morphology are often considered to be the result of ongoing interspecific hybridization; however, this conclusion is difficult to prove without analysis of chromosomal and/or molecular markers. In the butterfly genus Melitaea, such an intermediacy can be detected in male genitalia, and is more or less regularly observed in localities where two closely related, presumably parental species are found in sympatry. Here I analyze a high altitude Melitaea population from Mt. Hermon in north Israel and show that its male genitalia are clearly differentiated from those found in phenotypically similar M. persea and M. didyma, but in some aspects intermediate between them. This hybrid-like population is unique because, although M. didyma is present on Mt. Hermon, the true, low-altitude M. persea has never been reported from Israel. Cytogenetic analysis revealed no apomorphic chromosomal characters to distinguish the Mt. Hermon population from other known taxa of the M. persea and M. didyma species groups. At the same time, DNA barcode-based phylogeographic study showed that this population is ancient. It was estimated to originate 1–1.6 million years ago in the Levantine refugium from a common ancestor with M. persea. Generally, the data obtained are incompatible with interpretation of the studied population as a taxon conspecific with M. persea or M. didyma, or a swarm of recent hybrids between M. persea and M. didyma, although the possibility of ancient homoploid hybrid speciation cannot be ruled out. I also argue that the name Melitaeamontium assigned to butterflies from north Lebanon cannot be applied to the studied taxon from Mt. Hermon. Here I describe this morphologically and ecologically distinct entity as a new species Melitaeaacentria sp. n., and compare it with other taxa of the M.persea complex.
Holotype: (Fig. 2a, b), male, BOLD process ID BPAL2191-13, field # CCDB-17949_A06, GenBank accession number # KY777529; Israel, Mt. Hermon, 2050 m, 01 June 2013, A. Novikova leg., deposited in the Zoological Institute of the Russian Academy of Science (St. Petersburg).
Figure 3. Melitaea acentria in nature. Female. Israel, Mt. Hermon, 1800 m, 07 May 2016.
Figure 13. Melitaeaacentria and its habitat. Israel, Mt. Hermon, 2040 m, 22 June 2013.
Distribution: Melitaeaacentria is known to occur at high altitudes (1730–2060 m above the sea level) of Mt. Hermon (Fig. 11). Within these altitudes it is sympatric and syntopic with M. trivia syriaca, M. deserticola and M. cinxia. At the altitudes 1730–1780 m there is an essential overlapping of the M. acentria and M. didyma liliputana ranges where both species were found to fly together in early May 2016. Two other Melitaea species known from Mt. Hermon, M. collina and M. telona, were found to fly mostly at lower altitudes 1000–1600 m.
Etymology: The name acentria is a noun of the feminine gender. This name originates from the Greek prefix “a” that means “not” and from the Latin word “centrum” (centre) derived from the Greek “κέντρον” (kentron, a sharp point). Acentria is the Internet nickname of Asya Novikova who collected the samples initiated this research.This name indicates also the peripheral position of the new species within the distribution range of the M. persea species complex.
Figure 2. Melitaeaacentria sp. n. and M. persea persea. a M. acentria sp. n., holotype, male, sample 17949_A06, Israel, Mt. Hermon; upperside b M. acentria sp. n., holotype, male, sample 17949_A06, Israel, Mt. Hermon; underside c M. acentria sp. n., paratype, male, sample 25453_E09, Israel, Mt. Hermon d M. acentria sp. n., paratype, female, sample 25453_E11, Israel, Mt. Hermon eM. perseapersea, male, 17966_A10, Iran, Fars prov., Fasa area, 20 km W Estahban, 2200 m, 9-11 May 2007, B. Denno coll., MGCL accession # 2010-20 fM. persea persea, female, 17951_B01, Iran, Fars prov., 20 km N Darab, 2100-2300 m, 24.05.1999, leg. P. Hofmann, MGCL gM. persea persea, male, 17966_A11, Iran, Fars prov., Fasa area, 20 km W Estahban, 2200 m, 9–11.05.2007, MGCL accession # 2008-43 hM. persea persea, male, 17951_B02, Iran, Char Mahall-o-Bahtiyari, Umg. Shahr-e-Kord, 2000 m, 28 May 2002, leg. P. Hofmann, MGCL.
Figure 10.Melitaea persea persea, presumptive hybrid between M. interrupta and M. persea, M. persea paphlagonia, M. higginsi, M. didyma liliputana and M. interrupta. aMelitaea perseapersea, female, 17951_B03, Iran, Esfahan, Kuh-e-Marsenan, near Zefre, 2000 m, 26 May 2002, leg. Hofmann, MGCL b presumptive hybrid female between M. interrupta and M. persea, 17966_F12, Armenia, Zhangezur Range, Megri district, Litchk, 1800 m, 23 July 1999, A. Dantchenko leg., MGCL c M. persea paphlagonia, male, 17951_F11, Iran, Khorasan, Kuh-e-Binalut, 15 km SW Zoshk, 2300–2500 m, 7 June 1999, leg. P. Hofmann, MGCL d M. persea paphlagonia, male, 17951_F11, Iran, Khorasan, Kuh-e-Binalut, 15 km SW Zoshk, 2300-2500 m, 7 June 1999, leg. P. Hofmann, MGCL e M.higginsi, male, 17966_A12, Afghanistan, Hindukush, Panchir Valley, 20 June 2004, M.J.Simon collection, MGCL f M. higginsi, female, 17950_H10, Afghanistan, Badakhshan, Mt. Yamak N of Anjuman Pass, 3500-4000 m, 1-25 July 2004 M.J.Simon collection, MGCL g M. didyma liliputana, male, 17968_E10, Israel, Mt. Hermon h M. interrupta, male, 17966_F11, Armenia Armenia, Zhangezur Range, Kadjaran, 2500 m, 21–22 July 1999, leg. A. Chuvilin, MGCL; the wing underside is with black scales along the veins.
The Melitaeapersea species complex consists of the following taxa:
• M. persea Kollar, 1849
M. persea persea Kollar, 1849 (East Turkey, Armenia, Azerbaijan, Daghestan in Russian Caucasus, western, central and nothern parts of Iran)
M. persea paphlagonia Fruhstorfer, 1917 (NE Iran, probably also S. Turkmenistan)
• M. eberti Koçak, 1980 (N. Iran)
• M. higginsi Sakai, 1978 (Afghanistan)
• M. acentriaLukhtanov sp. n. (Mt. Hermon in Israel, definitely also the neighboring territories of Syria and Lebanon)
The identity and taxonomic status of the M. persea-similar samples from north Lebanon, Jordan, Iraq, Pakistan, and Afghanistan remain still unclear. The populations from Lebanon characterized by the mitochondrial haplogroup P2 (Fig. 9) could actually represent (i) a distinct subspecies of M. persea, (ii) an undescribed subspecies of M. acentria, or even (iii) an undescribed species. Further morphological, molecular and chromosomal studies are required to select between these hypotheses.
Vladimir A. Lukhtanov. 2017. A New Species of Melitaea from Israel, with Notes on Taxonomy, Cytogenetics, Phylogeography and Interspecific Hybridization in the Melitaeapersea complex (Lepidoptera, Nymphalidae). Comparative Cytogenetics. 11(2); 325-357. DOI: 10.3897/CompCytogen.v11i2.12370
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شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
Bufo (Anaxyrus) boreas species complex distribution. a) Bufo (Anaxyrus) boreas distribution (shown in brown) across the Western United States with hydrological Great Basin shown with black outline and hash mark interior; b) Bufo (Anaxyrus) boreas species complex and ranges for toads including new species, illustrating the narrow distribution of localized endemics. Spatial data for all toads except B. williamsi provided by IUCN (2015).
Images taken by M.R.Gordon exceptB. canorus with photo credit to G. Nafis.
We describe a new species of toad from the Great Basin region of northern Nevada belonging to the Bufo (Anaxyrus) boreas species complex. This cryptic species was detected through genetic analyses of toad populations sampled throughout the Great Basin and the morphological evidence was quantified through extensive sampling of live toads within the region. The new species has the smallest body size in the species complex, and can be further diagnosed from other species in the complex by its large tibial glands and unique coloration. The known distribution of the new species is restricted to an area less than 6 km2 in Dixie Valley, Churchill Co., Nevada. The Great Basin is an arid region where aquatic resources are both rare and widely scattered, making habitat suitable for anuran populations highly vulnerable to anthropogenic change. The habitat occupied by this newly described species is threatened by the incipient installation of geothermal and solar power development projects that require the water that defines its habitat.
Keywords: Amphibia, Bufo (Anaxyrus) williamsi sp. nov., Dixie Valley Toad, Western Toad, Bufo(Anaxyrus) boreas species complex, cryptic species, morphology, new species, conservation, geothermal
FIGURE 4. Photographs of Bufo (Anaxyrus) williamsi sp. nov. holotype (CAS 259271). Adult male presented live: (a) dorsal view and (b) ventral view; and preserved: (c) dorsal view and (d) ventral view.
Photographs taken by M.R.Gordon.
FIGURE 2. Bufo (Anaxyrus) boreas species complex distribution. a) Bufo (Anaxyrus) boreas distribution (shown in brown) across the Western United States with hydrological Great Basin shown with black outline and hash mark interior; b) Bufo (Anaxyrus) boreas species complex and ranges for toads including new species, illustrating the narrow distribution of localized endemics. Spatial data for all toads except B. williamsi provided by IUCN (2015).
Images taken by M.R.Gordon except B. canorus with photo credit to G. Nafis.
Bufo(Anaxyrus) williamsi sp. nov.
Dixie Valley Toad
Diagnosis. Bufo (Anaxyrus) williamsi is distinguishable from B. boreas by a combination of diagnostic morphological characters (Fig. 4; Table 1, Table 2), genetic evidence (Fig.3, Fig. 6), and localized distribution (Fig. 2b). Bufo (Anaxyrus) williamsi is distinct from B. boreas by: a small adult body size (SVL is more than 2.5 cm smaller than B. boreas; Table 1); significantly, but modestly, larger, closely-set eyes, and smaller head (Table 2); statistically and perceptibly larger tympanum, and shorter hind limbs; conspicuously large and elevated tibial glands; and distinctive color pattern (Fig. 4a, Fig. 4b).
FIGURE 2.Bufo (Anaxyrus) boreas species complex distribution. a) Bufo (Anaxyrus) boreas distribution (shown in brown) across the Western United States with hydrological Great Basin shown with black outline and hash mark interior; b) Bufo (Anaxyrus) boreas species complex and ranges for toads including new species, illustrating the narrow distribution of localized endemics. Spatial data for all toads except B. williamsi provided by IUCN (2015).
Images taken by M.R.Gordon except B. canorus with photo credit to G. Nafis.
Etymology. The specific epithet is in tribute to Robert Williams, former Field Supervisor of the U.S. Fish and Wildlife, whose Herculean efforts on behalf of the fauna of Nevada and California were critically important in discovering additional biodiversity of anurans in the Great Basin, and in focusing on the needs to provide protection to the rare and imperiled fauna, and the ecosystems upon which they depend, in Nevada and California. The Dixie Valley toad would not have been discovered without the efforts of this courageous public servant.
Distribution. Bufo (Anaxyrus) williamsi is found only within wetlands of limited extent fed from artesian springs on the western edge of the Dixie Valley Playa, east of the Stillwater Range in Dixie Valley, NV (Fig. 2b).
Natural history.Bufo (Anaxyrus) williamsi is restricted to the spring fed-wetland habitat along the western edge of the Dixie Valley playa. Similar to other toads in the B. boreas complex (except perhaps B. exsul, which is more aquatic), the terrestrial B. williamsi is typically nocturnal, emerging at dusk, and can be found in moist vegetation or in very still, shallow water with very little vegetation canopy. Dixie Valley experiences extreme temperature fluctuations between day and night temperatures, as well as season-to-season extremes, characteristic of cold desert ecosystems.
Michelle R. Gordon, Eric T. Simandle and C. R. Tracy. 2017. A Diamond in the Rough Desert Shrublands of the Great Basin in the Western United States: A New Cryptic Toad Species (Amphibia: Bufonidae: Bufo (Anaxyrus)) discovered in Northern Nevada. Zootaxa. 4290(1); 123–139. DOI: 10.11646/zootaxa.4290.1.7
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
The Paracanthocobitis zonalternans species complex is revised based on analysis of morphological and molecular data. Three new species, P. nigrolineata, P. marmorata, and P. triangula are described, and P. phuketensis is removed from synonymy. All species are described morphologically, geographic ranges are delimited, and relationships are discussed for those for which molecular data (cytochrome c oxidase subunit 1 - COI) are available. In view of the morphological similarities of some of the species, a surprising result of this study was the moderately large genetic distances among species. Uncorrected COI p-distances between geographic clades of P. zonalternans ranged from 7.6–9.3%, suggesting that the species are reproductively isolated from one another even though morphological changes are minor. Paracanthocobitis phuketensis, distributed in several rivers draining to the Gulf of Thailand and to the Andaman Sea, shows considerable intraspecific variation that should be explored in detail for historical and ecological explanations. Cobitis chlorosoma McClelland, 1839 from Assam, India, is a synonym of P. botia, not P. zonalternans.
Cobitis zonalternans Blyth, 1860:172. Type locality: Tenasserim provinces, Burma. Two syntypes (Day, 1869:551; Day, 1872:186), presumed lost (Hora, 1929:319). Neotype, ZSM 27468, designated by Kottelat, 1990:42, Huai Mae Charno, 4 km south Amphoe Mae Ramat on road 1085 [approximately 16.967N, 98.567E], Tak Province, Salween drainage, Thailand.
Diagnosis. Paracanthocobitis zonalternans is distinguished from all other species of Paracanthocobitis by combination of incomplete lateral line ending near dorsal-fin insertion; 9½–10½ branched dorsal-fin rays; axillary pelvic lobe present; series of black blotches, usually without black stripe but with faint dusky stripe, along side of body, usually ending near dorsal-fin insertion; circular black spot on upper margin of caudal-fin base; black pigment extending from lateral stripe onto pectoral-fin base; no marmorated pattern anterior to dorsal-fin origin; dorsal saddles short, rarely connecting to lateral blotches; 8 (occasionally 7) branched upper caudal-fin rays.
Distribution.Paracanthocobitis zonalternans is known from the Salween River drainage in Thailand and Myanmar (Fig. 5).
Paracanthocobitis phuketensis (Klausewitz, 1957)
Phuket Zipper Loach
Noemacheilus phuketensis Klausewitz, 1957:195, fig. 1. Type locality: Phuket Island, Thailand. Holotype: SMF 3966.
Diagnosis. Paracanthocobitis phuketensis is distinguished from all other species of Paracanthocobitis by combination of incomplete lateral line ending near dorsal-fin insertion; 9½–10½ branched dorsal-fin rays; axillary pelvic lobe usually absent, occasionally rudimentary; series of black blotches, sometimes overlain with faint dusky stripe along side of body not obscuring lateral blotches, ending near caudal-fin; circular black spot on upper margin of caudal-fin base; black pigment not extending from black blotches along side of body onto pectoral-fin base; no marmorated pattern anterior to dorsal-fin origin; dorsal saddles longer than interspaces, frequently connecting to lateral blotches; 7–8 branched upper caudal-fin rays.
Distribution. Paracanthocobitis phuketensis is known from Peninsular Thailand in the Panang Tak drainage and on Phuket Island, south to the Perlis, Padang Sarai, and Ketil drainages of Peninsular Malaysia (Fig. 5).
Paracanthocobitis nigrolineata, new species Blacklined Zipper Loach Diagnosis.Paracanthocobitis nigrolineata is distinguished from all other species of Paracanthocobitis by combination of incomplete lateral line ending near dorsal-fin insertion; 9½–10½ branched dorsal-fin rays; axillary pelvic lobe present; black stripe along side of body, ending near caudal-fin, usually overlying black lateral blotches extending ventrally from lateral stripe; circular black spot on upper margin of caudal-fin base; black pigment extending from lateral stripe onto pectoral-fin base; no marmorated pattern anterior to dorsal-fin origin; black dorsal saddles short, not connecting to lateral blotches; 8 branched upper caudal-fin rays. Distribution. Paracanthocobitis nigrolineata is known from Irrawaddy and Sittang drainages of Myanmar and the Mae Khlong drainage of Thailand with a large geographic gap in-between (Fig. 5). This gap is likely due to the paucity of sampling in Myanmar. Etymology. The epithet nigrolineata refers to the black stripe along the side of the body. Paracanthocobitis triangula, new species Wedged Zipper Loach Diagnosis. Paracanthocobitis triangula is distinguished from all other species of Paracanthocobitis by combination of incomplete lateral line ending near dorsal-fin insertion; 9½–10½ branched dorsal-fin rays; axillary pelvic lobe present; series of black blotches, sometimes overlain with faint dusky stripe along side of body not obscuring lateral blotches, ending just beyond dorsal-fin insertion; small black triangular blotch in ocellus on upper margin of caudal-fin base; black pigment of the midlateral stripe not extending onto pectoral-fin base; no marmorated pattern between dorsal saddles and lateral blotches; dorsal saddles usually extending ventrally just past faint lateral stripe, usually connecting to lateral blotches; 8 branched upper caudal-fin rays. Distribution.Paracanthocobitis triangula is known from the Brahmaputra (Jamuna) and Meghna drainages of Bangladesh (Fig. 5). Etymology. The epithet triangula refers to the usual presence of a small black triangular blotch in the ocellus on the upper margin of the caudal peduncle.
Paracanthocobitis marmorata, new species Marmorated Zipper Loach
Diagnosis.Paracanthocobitis marmorata is distinguished from all other species of Paracanthocobitis by combination of incomplete lateral line ending just beyond dorsal-fin insertion; 9½–10½ branched dorsal-fin rays; no axillary pelvic lobe; no black stripe along side of body; marmorated pattern of dark lines and blotches between irregularly shaped dorsal saddles dark blotches along side of body; teardrop-shaped black spot on upper margin of caudal-fin base; 8 branched upper caudal-fin rays. Distribution.Paracanthocobitis marmorata is known from the Barak drainage of Assam, India (Fig. 5). Etymology. The epithet marmorata refers to the marmorated pattern on the nape and between the dorsal saddles and lateral blotches in lieu of the black stripe along the side of the body typical of other members of the P. zonalternans complex.
Randal A. Singer, John M. Pfeiffer and Lawrence M. Page. 2017. A Revision of the Paracanthocobitis zonalternans (Cypriniformes: Nemacheilidae) Species Complex with Descriptions of Three New Species. Zootaxa. 4324(1); 85–107. DOI: 10.11646/zootaxa.4324.1.5
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
The Rufescent Burrowing Frog, Fejervarya rufescens, is thought to have a wide distribution across the Western Ghats in Peninsular India. This locally abundant but secretive species has a short breeding period, making it a challenging subject for field studies. We sampled 16 populations of frogs morphologically similar to F. rufescens in order to understand the variation among populations found across the Western Ghats. Our study shows significant morphological and genetic differences among the sampled populations, suggesting that F. ‘rufescens’ is a complex of several undescribed species. Using evidence from morphology and genetics, we confirm the presence of five distinct species in this group and formally describe four as new. The new species were delineated using a phylogeny based on three mitochondrial genes (16S, COI and Cytb) and a haplotype network of a nuclear gene (Rag1). Hereafter, the distribution of F. rufescens is restricted to the state of Karnataka and adjoining regions of northern Kerala. Three new species (Fejervarya kadar sp. nov., Fejervarya manoharani sp. nov. and Fejervarya neilcoxi sp. nov.) are from regions south of Palghat gap in the state of Kerala, and one (Fejervarya cepfi sp. nov.) from the northern Western Ghats state of Maharashtra. These findings indicate that Fejervarya frogs of the Western Ghats are more diverse than currently known. Our results will also have implications on the conservation status of F. rufescens, which was previously categorized as Least Concern based on its presumed wide geographical distribution. Furthermore, in order to facilitate a better taxonomic understanding of this region’s fejervaryan frogs, we divide all the known Fejarvarya species of the Western Ghats into four major groups—Fejervarya nilagirica group, Fejervaryarufescens group, Fejervaryasahyadris group and Fejervarya syhadrensis group, based on their morphological affinities.
Keywords: Amphibians, bioacoustics, multi-gene DNA barcoding, India, integrative taxonomy, molecular phylogeny, new species, species diversity, Western Ghats
Taxonomic accounts and description of new species
Fejervarya rufescens (Jerdon, 1853)
Rufescent Burrowing Frog (Daniels 2005)
Original name: Pyxicephalus rufescens Jerdon, 1853. Catalogue of reptiles inhabiting the Peninsula of India, Journal of the Asiatic Society of Bengal, 22: 522–534.
Fejervarya kadar sp. nov.
Kadar Burrowing Frog
Etymology. The species is named after the Kadar tribe of Kerala, who live in the Vazhachal forest where the type series was collected. We enjoyed their support and hospitality during amphibian field studies in the region. The specific epithet kadar is treated as an invariable noun in apposition to the generic name.
Fejervarya manoharani sp. nov.
Manoharan’s Burrowing Frog
Etymology: This species is named for Mr TM Manoharan, who severed as the Head of Kerala Forest Department for over a decade, for providing encouragement as well as personal financial support to SDB during the initial phases of his scientific career. The species epithet manoharani is treated as a noun in the genitive case.
Fejervarya neilcoxi sp. nov.
Neil Cox’s Burrowing Frog
Etymology: This species is named for Dr Neil Cox, Manager of the IUCN-Conservation International Biodiversity Assessment Unit. Neil has been associated with the IUCN Red List in a variety of capacities including species assessment and management, and the new species is named particularly in appreciation of his contribution towards the Global Amphibian Assessment. The species epithet neilcoxi is treated as a noun in the genitive case.
Fejervarya cepfi sp. nov.
CEPF Burrowing Frog
Etymology. The species is named after the Critical Ecosystem Partnership Fund www.cepf.net (CEPF) for its effort to protect global biodiversity hotspots by providing grants in general, and specifically for a grant supporting research and conservation planning in the Western Ghats biodiversity hotspot through the Project Western Ghats Network of Protected Areas for Threatened Amphibians www.wnpata.org (WNPATA) to SDB (University of Delhi). The specific epithet cepfi is treated as a noun in the genitive case.
Sonali Garg and S.D. Biju. 2017. Description of Four New Species of Burrowing Frogs in the Fejervarya rufescens complex (Dicroglossidae) with notes on Morphological Affinities of Fejervarya Species in the Western Ghats.
S K. Kiran, V. S. Anoop, K. C. Sivakumar, Raghunathan Dinesh, J. P. Mano, Deuti Kaushik and George Sanil. 2017. An Additional Record of Fejervarya manoharani Garg and Biju from the Western Ghats with A Description of Its Complete Mitochondrial Genome. Zootaxa. 4277(4); 491–502. DOI: 10.11646/zootaxa.4277.4.2
كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا
شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا
رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا
تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا
ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.
