[Paleontology • 2017] Corythoraptor jacobsi • High Diversity of the Ganzhou Oviraptorid Fauna Increased by A New “Cassowary-like” Crested Species ---ScRaBBlE

Corythoraptor jacobsi

Lü, Li, Kundrát, Lee, Sun, Kobayashi, Shen, Teng & Liu, 2017

  DOI: 10.1038/s41598-017-05016-6 

Abstract

A new oviraptorid dinosaur from the Late Cretaceous of Ganzhou, bringing oviraptrotid diversity of this region to seven taxa, is described. It is characterized by a distinct cassowary-like crest on the skull, no pleurocoels on the centra from the second through fourth cervical vertebrae, a neck twice as long as the dorsal vertebral column and slightly longer than the forelimb (including the manus). Phylogenetic analysis recovers the new oviraptorid taxon, Corythoraptor jacobsi, as closely related to Huanansaurus from Ganzhou. Osteochronology suggests that the type specimen of Corythoraptor had not reached stationary growth stage but died while decreasing growth rates. The histology implies that it would correspond to an immature individual approximately eight years old. We hypothesize, based on the inner structure compared to that in modern cassowaries, that the prominent casque of Corythoraptor was a multifunction-structure utilized in display, communication and probably expression of the fitness during mating seasons.

Figure 1: The holotype of Corythoraptor jacobsi gen. et sp. nov. (JPM-2015-001). (a) Photograph. (b) Outline drawings. (c) Close up of the skull and lower jaw, showing the pneumatic cassowary-like crest (Only skull and lower jaw elements are labeled). (d) Skeletal reconstruction (missing parts are in grey).

 Abbreviations: aof, antorbital fenestra; cav. caudal vertebrae; cr. cervical ribs; cv. cervical vertebrae; dv. dorsal vertebrae; fe, femur; fi. fibula; h, humerus; il, ilium; is, ischium; l, lacrimal; lj, lower jaw; ltf, lower temporal fenestra; m, maxilla; n, nasal; nar, narial opening; o, orbit; oc, occipital condyle; p, parietal; pm, premaxilla; po, postorbital; ps, pes; psc, pneumatic skull crest; pu. pubis; q, quadrate; ra, radius; sk, skull; sq, squamosal; stf, super temporal fenestra; ti, tibia; ul, ulna. Scale bar = 8 cm in (c) and 100 cm in (d).

Systematic palaeontology

Oviraptorosauria Barsbold, 1976.

Oviraptoridae Barsbold, 1976.

Corythoraptor jacobsi gen. et sp. nov.

Etymology: The generic name Corythoraptor refers to a raptor bearing a “cassowary-like crest” on its head, and the specific name is in honor of Professor Louis L. Jacobs, who has contributed to dinosaur research and has given excellent mentoring to three authors (JLü, YL and YK) when they were Ph.D. students at Southern Methodist University, Dallas, Texas, USA.

Type locality and horizon: A site in the vicinity of the Ganzhou Railway Station (GPS coordinates are provided on request from the first author), Ganzhou City; Campanian-Maastrichtian; Nanxiong Formation (Upper Cretaceous).

 Diagnosis: An oviraptorosaurian dinosaur with the following unique combination of characters: ratio of the length of the tomial margin of the premaxilla to the premaxilla height (ventral to the external naris) is 1.0–1.4; inclination of the anteroventral margin of the premaxilla relative to the horizontally positioned ventral margin of the jugal posterodorsal; antorbital fossa bordered anteriorly by the maxilla; narial opening much longer than width; infratemporal fenestra dorsoventrally elongate, narrow anteroposteriorly; the supranarial process of the premaxilla bears two processes: a short posterodorsally extending process, forming the anterodorsal margin of the external nasal opening, and a long process, forming most of the anterodorsal process of the premaxilla; distinct cassowary-like helmet on the skull; long axis of the external narial opening parallel to the dorsal margin of antorbital fenestra; straight anterodorsal margin of dentary in lateral view; a deep fossa, sometimes with associated pneumatopore on lateral surface of dentary; no pleurocoels on the centra from the second through fourth cervical vertebrae; the length of the neck twice as long as the dorsal vertebral column, and slightly longer than the entire forelimb length (including the manus); less pronounced deltopectoral crest of humerus, forming an arc rather than being quadrangular; ratio of the length of the manus to the length of the humerus plus the radius between 0.50 and 0.65; the ungual of digit III less curved than other unguals; lesser trochanter (cranial trochanter) completely fused with the greater trochanter and distal ends of shafts of metatarsal II straight and metatarsal IV laterally deflected. 

Corythoraptor jacobsi gen. et sp. nov. is assigned to oviraptorid dinosaurs based on the following characters: proximal caudals with pneumatized centra; ischium with its posterior profile concave3; premaxilla pneumatized; the subantorbital portion of the maxilla inset medially; the palate extending below the cheek margin; the external naris overlapping most of the antorbital fossa rostrodorsally; the bones of the skull roof pneumatized; the pubic shaft concave cranially, the mandibular symphysis tightly sutured; the shortened preorbital region, and the toothless jaws.

.....

Figure 2: The cranial casque of Corythoraptor jacobsi and recent cassowaries. (a–c) the crested skull of Corythoraptor and head appearance restorations. (d) a close-up (see dotted rectangle in a and b) of eroded bony shell in the posterolateral casque of Corythoraptor. (e) the crested skull of the recent cassowary (Casuarius uniappendiculatus; Museum für Naturkunde in Berlin, Germany: MfN-ZMB 93274). (f) a keratinous helmet over the skull of the recent cassowary (unnumbered specimen of Casuarius casuarius from the osteological collections of ZOO Protivín, Czech Republic). (g,h) coronal cuts through the cassowary skull – (g) Casuarius casuarius MfN-ZMB 36820, (h) Casuarius casuarius: MfN-ZMB 36885 (see dotted lines in f); note transition in strut-like trabecular arrangement. (i) close-up to contact between keratinous and skeletal components of the casque in recent cassowary, unnumbered specimen of Casuarius sp. from the osteological collections of Field Museum of Natural History, Chicago, USA.

Abbreviations: cq, casque; cr, cranium; exs, external surface; kesh, keratinous sheath; or, orbit; tr, trabeculae.

Figure 5: The living scene of Corythoraptor jacobsi gen. et sp. nov.

Drawn by Zhao Chuang

 

Junchang Lü, Guoqing Li, Martin Kundrát, Yuong-Nam Lee, Zhenyuan Sun, Yoshitsugu Kobayashi, Caizhi Shen, Fangfang Teng and Hanfeng Liu. 2017. High Diversity of the Ganzhou Oviraptorid Fauna Increased by A New “Cassowary-like” Crested Species.
  Scientific Reports. 7, Article number: 6393. DOI: 10.1038/s41598-017-05016-6

ResearchGate.net/publication/318726539_High_diversity_of_the_Ganzhou_Oviraptorid_Fauna_increased_by_a_new_cassowary-like_crested_species

Introducing Corythoraptor jacobsi.  paleonerdish.wordpress.com/2017/07/28/introducing-corythoraptor-jacobsi/ via @ferwen

Paleo Profile: The Crested Thief  blogs.scientificamerican.com/laelaps/paleo-profile-the-crested-thief/

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

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شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Beibeilong sinensis • Perinate and Eggs of A Giant Caenagnathid Dinosaur from the Late Cretaceous of central China ---ScRaBBlE

 Beibeilong sinensis

Pu, Zelenitsky, Lü, Currie, Carpenter, Xu, Koppelhus, Jia, Xiao, Chuang, Li, Kundrát & Shen, 2017DOI: 10.1038/NComms14952

Abstract

The abundance of dinosaur eggs in Upper Cretaceous strata of Henan Province, China led to the collection and export of countless such fossils. One of these specimens, recently repatriated to China, is a partial clutch of large dinosaur eggs (Macroelongatoolithus) with a closely associated small theropod skeleton. Here we identify the specimen as an embryo and eggs of a new, large caenagnathid oviraptorosaur, Beibeilong sinensis. This specimen is the first known association between skeletal remains and eggs of caenagnathids. Caenagnathids and oviraptorids share similarities in their eggs and clutches, although the eggs of Beibeilong are significantly larger than those of oviraptorids and indicate an adult body size comparable to a gigantic caenagnathid. An abundance of Macroelongatoolithus eggs reported from Asia and North America contrasts with the dearth of giant caenagnathid skeletal remains. Regardless, the large caenagnathid-Macroelongatoolithus association revealed here suggests these dinosaurs were relatively common during the early Late Cretaceous.

Systematic palaeontology 

Theropoda Marsh 1881

Oviraptorosauria Barsbold 1976

Caenagnathidae Sternberg 1940

 Beibeilong sinensis gen. et sp. nov.

Figure 1: Photograph of eggs and skeleton of Beibeilong sinensis (HGM 41HIII1219).
Right image shows schematic overlay of approximate locations of individual eggs. Eggs 1 through 4 are in an upper layer just beneath the skeleton, whereas Egg 5 is in a lower layer of the block. Scale bar is in centimetre.

The fossil (left) and a digital overlay (right) shows where the eggs were placed before they broke and fossilized. Notice the skeleton of the embryo next to the ruler. 

Holotype. HGM 41HIII1219, a small, semi-articulated skeleton (‘Baby Louie’) associated with a partial nest of 6–8 eggs. The specimen is housed in the Henan Geological Museum (HGM), Zhengzhou, China.

the curled embryo of Beibelong on top of the eggs (eggshell is dark grey in color).

Photograph by Darla Zelenitsky, University of Calgary 

An illustration of Beibeilong embryo and egg fossil, with reconstruction of skeleton and fleshed-out models to the right. 

Figure 5: Reconstruction of Beibeilong embryo in ovo.
The drawing shows the approximate size of the Beibeilong embryo inside a Macroelongatoolithus egg.

drawn by Vladimir Rimbala

 Diagnosis. A large caenagnathid that has the following unique suite of features: antorbital fossa demarcated by sharply defined alveolar and dorsoposterior trending ridges, posterodorsal margin of lacrimal overlapped by frontal, subantorbital portion of the maxilla is inset medially, pronounced retroarticular process with a distinct concave posterior facet (roughly as tall at the base as it is wide), preacetabular process of the ilium longer than postacetabular process, posterior end of the postacetabular process truncated or broadly rounded, and accessory trochanter of the femur weakly developed.

 Locality and horizon. The specimen was discovered at a latitude/longitude of 33°15′30″ N, 111°43′41″ E in Heimaogou, 2 km east of Zhaoying Village, Yangcheng Township, Xixia County, Henan Province, People’s Republic of China. The locality is in the Upper Cretaceous Gaogou Formation (Cenomanian—Turonian)

Etymology. The generic name is derived from Chinese Pinyin ‘beibei’ for baby and ‘long’ for dragon. The specific name is derived from Latin referring to its discovery in China.

Beibeilong sinensis lived about 90 million years ago, during the Cretaceous period. The adults of this species were enormous, measuring up to 8 meters long from the snout to the end of the tail, and weighing up to 3,000 kilograms when fully grown at age 11.

Illustration: Zhao Chuang  

  

Hanyong Pu, Darla K. Zelenitsky, Junchang Lü, Philip J. Currie, Kenneth Carpenter, Li Xu, Eva B. Koppelhus, Songhai Jia, Le Xiao, Huali Chuang, Tianran Li, Martin Kundrát and Caizhi Shen. 2017. Perinate and Eggs of A Giant Caenagnathid Dinosaur from the Late Cretaceous of central China.  Nature Communications. 8, Article number: 14952. DOI: 10.1038/ncomms14952

First baby of a gigantic Oviraptor-like dinosaur belongs to a new species

 eurekalert.org/e/7Xft via @UCalgary @EurekAlert

'Baby Dragon' Dinosaur Found Inside Giant Egg  on.NatFeo.com/2pYHLPO via @NatGeo

Embryo of Colossal Dinosaur Was Preserved for 90 Million Years

shar.es/1FDLTy via @LiveScience

  

   

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2019] Gobiraptor minutus • A New Baby Oviraptorid Dinosaur (Dinosauria: Theropoda) from the Upper Cretaceous Nemegt Formation of Mongolia ---ScRaBBlE

Gobiraptor minutus 

Lee, Lee, Chinsamy, Lü, Barsbold & Tsogtbaatar, 2019

 DOI: 10.1371/journal.pone.0210867

Abstract

Recent discoveries of new oviraptorosaurs revealed their high diversity from the Cretaceous Period in Asia and North America. Particularly, at the family level, oviraptorids are among the most diverse theropod dinosaurs in the Late Cretaceous of Mongolia and China. A new oviraptorid dinosaur Gobiraptor minutus gen. et sp. nov. from the Upper Cretaceous Nemegt Formation is described here based on a single holotype specimen that includes incomplete cranial and postcranial elements. The most prominent characters of Gobiraptor are its thickened rostrodorsal end of the mandibular symphysis and a rudimentary lingual shelf on each side of the dentary. Each lingual shelf is lined with small occlusal foramina and demarcated by a weakly developed lingual ridge. This mandibular morphology of Gobiraptor is unique among oviraptorids and likely to be linked to a specialized diet that probably included hard materials, such as seeds or bivalves. The osteohistology of the femur of the holotype specimen indicates that the individual was fairly young at the time of its death. Phylogenetic analysis recovers Gobiraptor as a derived oviraptorid close to three taxa from the Ganzhou region in southern China, but rather distantly related to other Nemegt oviraptorids which, as the results of recent studies, are also not closely related to each other. Gobiraptor increases diversity of oviraptorids in the Nemegt Formation and its presence confirms the successful adaptation of oviraptorids to a mesic environment.

Fig 4. Postcranial elements of the holotype specimen (MPC-D 102/111) of Gobiraptor minutus gen. et sp. nov.
 (A) Skeletal reconstruction in left lateral view (missing and damaged portions of the bones in gray). (B) Left ilium in lateral view. (C) Proximal caudal vertebrae in left lateral view with close-up of the infraprezygapophyses. (D) Chevron in cranial view. (E-F) Right scapula in dorsal (E) and lateral (F) views. (G) Last sacral and the two proximalmost caudals in left lateral view. (H) Right pubis in medial view. (I) Right ischium in lateral view. (J) Right femur in distal view. (K) Left metatarsus and distal tarsals in proximal view. (L) Right femur in cranial view. (M-N) Left metatarsus in lateral (M) and dorsal (N) views. 

Abbreviations: acr, acromion process; ant, antitrochanter; ch, chevron; cv, caudal vertebra(e); diprf, dorsal infraprezygapophyseal fossa; dt, distal tarsal(s); fct, cranial trochanter of femur; fh, femoral head; gl, glenoid fossa; idf, infradiapophyseal fossa; lc, lateral condyle; mc, medial condyle; mep, medial epicondyle; miprf, middle infraprezygapophyseal fossa; mt II, metatarsal II; mt IV, metatarsal IV; mt V, metatarsal V; ns, neural spine; obp, obturator process; pra, preacetabular process; pup, pubic peduncle; sprf, supraprezygapophyseal fossa; sv, sacral vertebra; tfc, tibiofibular crest; tp, transverse process; viprf, ventral infraprezygapophyseal fossa. Scale bars equal 10 cm in (A); 1 cm in (B-N).

Systematic paleontology

Dinosauria Owen, 1842

Theropoda Marsh, 1881

Maniraptora Gauthier, 1986

Oviraptorosauria Barsbold, 1976

Oviraptoridae Barsbold, 1976

Gobiraptor minutus gen. et sp. nov.

Etymology: The generic name Gobiraptor is a combination of ‘Gobi’ which refers to the Gobi Desert where the holotype specimen was found and ‘raptor’ which is Latin for thief. The specific name ‘minutus’ is Latin for small and refers to the small size of the holotype specimen.

Holotype: The holotype specimen (MPC-D 102/111) (Figs 2–4, S2 and S3 Figs) consists of mostly incomplete cranial and postcranial elements including ventral parts of the premaxillae and maxillae, a tip of the right jugal, fused vomer, parts of articulated pterygoids and ectopterygoids, incomplete right palatine, central part of the left postorbital, partial right quadrate and quadratojugal, incomplete lower jaw, with most of its elements broken, the last sacral vertebra which is articulated with the two proximalmost caudal vertebrae, articulated but incomplete proximal caudal vertebrae, fragments of chevrons, partial right scapula and humerus, incomplete pelvic girdles, nearly complete both femora, complete left metatarsus with distal tarsals 3 and 4, incomplete left pedal digits I, III, and IV, and several unidentified fragments. MPC-D 102/111 was also found with other theropod skeletons including postcranial elements of alvarezsaurids and larger oviraptorids.

Type locality and horizon: Altan Uul III, Ömnögovi Province, Mongolia (Fig 1, S1 Fig). Upper Cretaceous Nemegt Formation.

Diagnosis: Gobiraptor minutus is an oviraptorid dinosaur diagnosed by the following unique set of characteristics (autapomorphies are marked with an asterisk): a flat articular surface for the quadratojugal on the quadrate*; rostrocaudally elongate dentary rostral to the external mandibular fenestra; extremely thickened rostrodorsal end of the mandibular symphysis with a caudal expansion of its dorsal surface *; a rudimentary lingual triturating shelf on each dentary bearing small occlusal foramina*; a weakly developed lingual ridge on each lingual shelf*; absence of any prominent symphyseal ventral process of the dentary; coronoid bone present; the rostral end of the coronoid bone wedging into the ventral surface of the dorsal ramus of the dentary*; cranial trochanter of the femur separated from the greater trochanter with a distinct furrow between them.

....

   

Conclusions: 

Gobiraptor minutus gen. et sp. nov. is a new derived oviraptorid represented by an incomplete skeleton including both cranial and postcranial elements. Gobiraptor is primarily distinguished from other oviraptorids by its dentary with the extremely thickened rostrodorsal end of the mandibular symphysis, lingual ridges and lingual shelves bearing occlusal foramina. The unique morphology of the mandible of Gobiraptor is probably closely related to a crushing-related feeding style and a specialized diet, which may have incorporated hard seeds or shelled organisms. Although Gobiraptor was recovered from the Nemegt Formation, its phylogenetic position showed a close relationship with three Ganzhou oviraptorids. The distant relationships among the Nemegt oviraptorids on the phylogenetic tree were reaffirmed in this study. Therefore, it is highly unlikely that the evolution of these unusually diverse animals was facilitated by a simple sympatric speciation. The presence of Gobiraptor in the Nemegt Formation, together with occurrences of other oviraptorids, also indicates that abundant oviraptorids lived in mesic environments and they were one of the most diverse and successful groups of dinosaurs in the Nemegt region.

      

 Sungjin Lee, Yuong-Nam Lee , Anusuya Chinsamy, Junchang Lü†, Rinchen Barsbold and Khishigjav Tsogtbaatar. 2019. A New Baby Oviraptorid Dinosaur (Dinosauria: Theropoda) from the Upper Cretaceous Nemegt Formation of Mongolia.  PLoS ONE. 14(2): e0210867. DOI: 10.1371/journal.pone.0210867

New oviraptorosaur species discovered in Mongolia phys.org/news/2019-02-oviraptorosaur-species-mongolia.html via @physorg_com

     

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2018] Anomalipes zhaoi • A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Comments on Size Variation Among Oviraptorosaurs ---ScRaBBlE

Anomalipes zhaoi 

 Yu, Wang, Chen, Sullivan, Wang, Wang & Xu, 2018

  DOI: 10.1038/s41598-018-23252-2 

Abstract

The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi differs from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with significant posterior orientation; accessory trochanter low and confluent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior flange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae.

Figure 2 Preserved left femur, tibia, and fibula of Anomalipes zhaoi ZCDM V0020.
 Left femur in anterior (a), posterior (b), lateral (c), medial (d) and proximal (e) views.
Left tibia in anterior (f), posterior (g) and distal (j) views. Shading indicates the articular facet for the ascending process of the astragalus.
Left fibula in lateral (h) and medial (i) views.

Abbreviations: act, accessory trochanter; dg, distinct groove; fc, fibular crest; fh, femoral head; ft, fourth trochanter; gt, greater trochanter; if, iliofibularis tubercle; ig, intercondylar groove; lm, lateral malleolus; lr, lateral ridge; lt, lesser trochanter; mm, medial malleolus; pt, posterior trochanter; taf, triangular articular facet. Scale bar 1 cm.

Figure 3 Preserved pedal elements of Anomalipes zhaoi ZCDM V0020.
Left metatarsal III in lateral (a), medial (b), posterior (c), anterior (d), proximal (e) and distal (f) views. Dark lines indicate ridges on the posterior surface of the shaft.
Phalanx IV-1 in lateral (g), medial (h), proximal (i), and distal (j) views. Phalanx II-3 in lateral (k) and medial (l) views.

Abbreviations: fl, flexor tubercle; lc, lateral condyle; lgf, ligament fossa; pdc, proximal dorsal crest; pdl, proximal dorsal lip; vr, ventral ridge; ptaf, proximal triangular articular facet; rlmh, ridge-like medial hemicondyle; vr, ventral ridge (extending to medial hemicondyle). Scale bar 1 cm.

Systematic palaeontology

Theropoda Marsh 1881

Oviraptorosauria Barsbold 1976

Caenagnathidae Sternberg 1940

Anomalipes zhaoi gen.et sp. nov

Etymology: Generic name is a combination of the Latin “Anomalus” and “pes”, referring to the unusual shape of the foot. Specific name is in honour of Xijin Zhao, a Chinese palaeontologist who has made great contributions to research on Zhucheng dinosaur fossils.

Holotype: ZCDM V0020 (Zhucheng Dinosaur Museum, Zhucheng, Shandong, China), an incomplete left hindlimb, including the left femur missing the distal end, the left tibia missing the proximal end, the left fibula missing the distal and proximal ends, a complete metatarsal III and two pedal phalanges. Although these bones are disarticulated, they are inferred to be derived from a single theropod individual given that 1) they were preserved in a small area of less than 0.3 square metres within a Shantungosaurus bonebed; and 2) no other theropod skeletal elements are preserved nearby.

Locality and horizon: Kugou, Zhucheng City, Shandong Province, China. Upper Cretaceous Wangshi Group.

Diagnosis: A new caenagnathid with the following unique combination of features: femoral head anteroposteriorly narrow and somewhat deflected posteriorly; accessory trochanter low; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior flange near proximal end, medial hemicondyle much narrower than lateral hemicondyle, and longitudinal groove on distal articular surface; and pedal phalanx II-3 with lateral collateral groove deeper and more dorsally located than medial groove.

Figure 5 Simplified oviraptorosaurian phylogenetic tree, showing size ranges for basal oviraptorosaurs, the Caenagnathidae, and the Oviraptoridae. Grey boxes represent body mass ranges for three oviraptorosaurian groups: basal oviraptorosaurs, oviraptorids, and caenagnathids. See the electronic supplementary material for estimated body masses of various oviraptorosaurian species.

 Abbreviations: Caud: Caudipteryx zoui; Avim: Avimimus portentosus; Conc: Conchoraptor gracilis; Micr: Microvenator celer; Wula: Wulatelong gobiensis; Citi: Citipati osmolskae; Nank: Nankangia jiangxiensis; Anom: Anomalipes zhaoi; Neme: Nemegtia barsboldi; Anzu: Anzu wyliei; Giga: Gigantoraptor erlianensis.

Yilun Yu, Kebai Wang, Shuqing Chen, Corwin Sullivan, Shuo Wang, Peiye Wang and Xing Xu. 2018. A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Comments on Size Variation Among Oviraptorosaurs. Scientific Reports. volume 8, Article number: 5030. DOI: 10.1038/s41598-018-23252-2 

 researchgate.net/publication/323961130_A_new_caenagnathid_dinosaur_from_the_Upper_Cretaceous_Wangshi_Group_of_Shandong_China

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

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من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2018] Incubation Behaviours of Oviraptorosaur Dinosaurs in Relation to Body Size ---ScRaBBlE

 egg incubation in oviraptorosaurs showing small species sat on the eggs (upper), whereas giant species rested in the central opening of the clutch (lower).

in Tanaka, Zelenitsky, Lü, et al., 2018. 

DOI: 10.1098/rsbl.2018.0135

Reconstruction by Masato Hattori.   marchan-forest.blogspot.com

Abstract
Most birds sit on their eggs during incubation, a behaviour that likely evolved among non-avian dinosaurs. Several ‘brooding' specimens of smaller species of oviraptorosaurs and troodontids reveal these non-avian theropods sat on their eggs, although little is known of incubation behaviour in larger theropod species. Here we examine egg clutches over a large body size range of oviraptorosaurs in order to understand the potential effect of body size on incubation behaviour. Eggshell porosity indicates that the eggs of all oviraptorosaurs were exposed in the nest, similar to brooding birds. Although all oviraptorosaur clutches consist of radially arranged eggs in a ring configuration, clutch morphology varies in that the central opening is small or absent in the smallest species, becomes significantly larger in larger species, and occupies most of the nest area in giant species. Our results suggest that the smallest oviraptorosaurs probably sat directly on the eggs, whereas with increasing body size more weight was likely carried by the central opening, reducing or eliminating the load on the eggs and still potentially allowing for some contact during incubation in giant species. This adaptation, not seen in birds, appears to remove the body size constraints of incubation behaviour in giant oviraptorosaurs.

KEYWORDS: dinosaur, egg, incubation, nest, Oviraptorosauria, Theropoda

 Reconstruction of egg incubation in oviraptorosaurs showing small species sat on the eggs (upper),
whereas giant species rested in the central opening of the clutch (lower).

Illustration is drawn by Masato Hattori.

Kohei Tanaka, Darla K. Zelenitsky, Junchang Lü, Christopher L. DeBuhr, Laiping Yi, Songhai Jia, Fang Ding, Mengli Xia, Di Liu, Caizhi Shen and Rongjun Chen. 2018. Incubation Behaviours of Oviraptorosaur Dinosaurs in Relation to Body Size. BIOLOGY LETTERS. DOI: 10.1098/rsbl.2018.0135

 marchan-forest.blogspot.com/2018/05/blog-post.html

    

How huge dinosaurs nested without crushing their eggs cbc.ca/1.4602808 

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

من هنا

او

من هنا

شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات

كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.