[Paleontology • 2017] Almas ukhaa • A New Late Cretaceous Troodontid from Ukhaa Tolgod, Ömnögovi Aimag, Mongolia ---ScRaBBlE

Almas ukhaa

Pei, Norell, Barta, Bever, Pittman & Xu, 2017

  DOI: 10.1206/3889.1 

ABSTRACT

A new troodontid dinosaur, Almas ukhaa, from the Late Cretaceous deposits of the Djadokhta Formation at Ukhaa Tolgod, Mongolia, is described here. The holotype specimen (IGM 100/1323) comprises an almost complete and articulated cranium and partial articulated postcranial skeleton. This specimen has a small body size and a short snout as in basal paravians, but it exhibits a number of derived troodontid features that differentiate Almas ukhaa from the Early Cretaceous troodontids reported from China and unite this new taxon with other Late Cretaceous troodontids. Relative to other troodontids, Almas ukhaa is autapomorphic in the presence of a posteriorly curved pterygoid flange, absence of a lateral groove on the anterior part of the dentary, presence of a distinct spikelike process on the ischium, and elongate chevrons. The eggshell associated with IGM 100/1323 can be assigned to Prismatoolithidae indet. based on the smooth surface, eggshell thickness, and microstructural characteristics, and also preserves attributes similar to Protoceratopsidovum minimum. A unique relationship between Byronosaurus and the perinate troodontids IGM 100/972 and IGM 100/974 is no longer supported based on the new observations of Almas ukhaa and Gobivenator.

FIGURE 2. Skull of IGM 100/1323 in right lateral view.

FIGURE 1. Type specimen of Almas ukhaa, IGM 100/1323.

SYSTEMATIC PALEONTOLOGY

THEROPODA MARSH, 1881

COELUROSAURIA HUENE, 1920

MANIRAPTORA GAUTHIER, 1986

TROODONTIDAE GILMORE, 1924

Almas ukhaa, new taxon

Etymology: Almas is in reference to the wild man or snowman of Mongolian mythology (Rincen, 1964). Ukhaa refers to the locality of Ukhaa Tolgod, discovered in 1993, where the specimen was collected. 

Rui Pei, Mark A. Norell, Daniel E. Barta, G.S. Bever, Michael Pittman and Xing Xu. 2017. Osteology of A New Late Cretaceous Troodontid Specimen from Ukhaa Tolgod, Ömnögovi Aimag, Mongolia. American Museum Novitates. 3889; 1-47.  DOI: 10.1206/3889.1
 digitallibrary.amnh.org/handle/2246/6818
ResearchGate.net/publication/321834802_Osteology_of_a_New_Late_Cretaceous_Troodontid_Specimen_from_Ukhaa_Tolgod_Omnogovi_Aimag_Mongolia

 twitter.com/Synapsida/status/941727444838465536

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

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شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2017] Zhongjianosaurus yangi • A New Tiny Dromaeosaurid Dinosaur from the Lower Cretaceous Jehol Group of western Liaoning and Niche Differentiation Among the Jehol Dromaeosaurids ---ScRaBBlE

Zhongjianosaurus yangi 

Xu & Qin, 2017 

Abstract 

The Early Cretaceous Jehol dromaeosaurids are taxonomically and morphologically diverse, and one of them, Microraptor zhaoianus, has been suggested to be among the smallest known non-avialan theropods. However, this idea is based on specimens of relatively early ontogenetic stages, and the lower limit of the mature body mass of Jehol dromaeosaurids thus remains unknown. Here we describe a new dromaeosaurid, Zhongjianosaurus yangi gen. et sp. nov., based on a specimen from the Lower Cretaceous Yixian Formation (the middle section of the Jehol Group) from Sihedang, Lingyuan County, Liaoning in Northeast China. While this new taxon is referable to the Microraptorinae, it differs from other microraptorine dromaeosaurids in numerous features, most notably the fusion of proportionally long uncinate processes to dorsal ribs, a humerus with a strongly medially offset proximal end and a large fenestra within the deltopectoral crest, an ulna slightly longer than the humerus, and an arctometatarsalian pes. Most significantly, the estimated 0.31 kg mass of the Z. yangi holotype of an adult individual confirms that some Jehol dromaeosaurids are among the smallest known non-avialan theropods. Our preliminary analysis demonstrates niche differentiation among the Jehol dromaeosaurids, a phenomenon rarely reported among Mesozoic dinosaurian faunas. 

Key words:  Lower Cretaceous, Jehol Group, Theropoda, Dromaeosauridae, small size, morphological variation, niche differentiation

 

Systematic paleontology

 Theropoda Marsh, 1881

 Tetanurae Gauthier, 1986

 Dromaeosauridae Matthew et Brown, 1922

Zhongjianosaurus yangi gen. et sp. nov.

 Etymology: The genus name and specific epithet are in honor of Yang Zhongjian (C.C. Young), who is the founder of vertebrate paleontology in China.

Locality and Horizon: Sihedang, Lingyuan County, Liaoning Province, China. Possibly Yixian Formation, Aptian, Cretaceous (Swisher et al., 2001).

Diagnosis: A tiny microraptorine theropod distinguishable from other microraptorines in the following autapomorphies: proportionally long ossified uncinate processes fused to dorsal ribs, widely arched furcula with slender and posteriorly curved clavicular rami, humeral proximal end strongly offset medially from humeral shaft, humeral internal tuberosity short, presence of large fenestra within humeral deltopectoral crest, humeral ulnar condyle hypertrophied, ulna slightly longer than humerus, ulnar olecranon process with posterior margin mediolaterally pinched, ulnar distal end bending anteriorly and strongly expanded laterally, proximal end of metacarpal II with strong ventrolateral extension, metacarpal III laterally bowed with longitudinal ventral groove, phalanx II-2 without proximodorsal lip and lacking strong dorsal arching, femoral head stout and lower than trochanteric crest, medial condyle of tibiotarsus distal end with a prominent distal extension, arctometatarsalian pes, and metatarsal II without ginglymus on distal end.

 

 Xing Xu and Zi-Chuan Qin. 2017. A New Tiny Dromaeosaurid Dinosaur from the Lower Cretaceous Jehol Group of western Liaoning and Niche Differentiation Among the Jehol Dromaeosaurids. Vertebrata PalAsiatica. In press.

http://www.ivpp.ac.cn/cbw/gjzdwxb/pressonline/201704/P020170410587218147108.pdf

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

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كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2018] Anomalipes zhaoi • A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Comments on Size Variation Among Oviraptorosaurs ---ScRaBBlE

Anomalipes zhaoi 

 Yu, Wang, Chen, Sullivan, Wang, Wang & Xu, 2018

  DOI: 10.1038/s41598-018-23252-2 

Abstract

The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi differs from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with significant posterior orientation; accessory trochanter low and confluent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior flange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae.

Figure 2 Preserved left femur, tibia, and fibula of Anomalipes zhaoi ZCDM V0020.
 Left femur in anterior (a), posterior (b), lateral (c), medial (d) and proximal (e) views.
Left tibia in anterior (f), posterior (g) and distal (j) views. Shading indicates the articular facet for the ascending process of the astragalus.
Left fibula in lateral (h) and medial (i) views.

Abbreviations: act, accessory trochanter; dg, distinct groove; fc, fibular crest; fh, femoral head; ft, fourth trochanter; gt, greater trochanter; if, iliofibularis tubercle; ig, intercondylar groove; lm, lateral malleolus; lr, lateral ridge; lt, lesser trochanter; mm, medial malleolus; pt, posterior trochanter; taf, triangular articular facet. Scale bar 1 cm.

Figure 3 Preserved pedal elements of Anomalipes zhaoi ZCDM V0020.
Left metatarsal III in lateral (a), medial (b), posterior (c), anterior (d), proximal (e) and distal (f) views. Dark lines indicate ridges on the posterior surface of the shaft.
Phalanx IV-1 in lateral (g), medial (h), proximal (i), and distal (j) views. Phalanx II-3 in lateral (k) and medial (l) views.

Abbreviations: fl, flexor tubercle; lc, lateral condyle; lgf, ligament fossa; pdc, proximal dorsal crest; pdl, proximal dorsal lip; vr, ventral ridge; ptaf, proximal triangular articular facet; rlmh, ridge-like medial hemicondyle; vr, ventral ridge (extending to medial hemicondyle). Scale bar 1 cm.

Systematic palaeontology

Theropoda Marsh 1881

Oviraptorosauria Barsbold 1976

Caenagnathidae Sternberg 1940

Anomalipes zhaoi gen.et sp. nov

Etymology: Generic name is a combination of the Latin “Anomalus” and “pes”, referring to the unusual shape of the foot. Specific name is in honour of Xijin Zhao, a Chinese palaeontologist who has made great contributions to research on Zhucheng dinosaur fossils.

Holotype: ZCDM V0020 (Zhucheng Dinosaur Museum, Zhucheng, Shandong, China), an incomplete left hindlimb, including the left femur missing the distal end, the left tibia missing the proximal end, the left fibula missing the distal and proximal ends, a complete metatarsal III and two pedal phalanges. Although these bones are disarticulated, they are inferred to be derived from a single theropod individual given that 1) they were preserved in a small area of less than 0.3 square metres within a Shantungosaurus bonebed; and 2) no other theropod skeletal elements are preserved nearby.

Locality and horizon: Kugou, Zhucheng City, Shandong Province, China. Upper Cretaceous Wangshi Group.

Diagnosis: A new caenagnathid with the following unique combination of features: femoral head anteroposteriorly narrow and somewhat deflected posteriorly; accessory trochanter low; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior flange near proximal end, medial hemicondyle much narrower than lateral hemicondyle, and longitudinal groove on distal articular surface; and pedal phalanx II-3 with lateral collateral groove deeper and more dorsally located than medial groove.

Figure 5 Simplified oviraptorosaurian phylogenetic tree, showing size ranges for basal oviraptorosaurs, the Caenagnathidae, and the Oviraptoridae. Grey boxes represent body mass ranges for three oviraptorosaurian groups: basal oviraptorosaurs, oviraptorids, and caenagnathids. See the electronic supplementary material for estimated body masses of various oviraptorosaurian species.

 Abbreviations: Caud: Caudipteryx zoui; Avim: Avimimus portentosus; Conc: Conchoraptor gracilis; Micr: Microvenator celer; Wula: Wulatelong gobiensis; Citi: Citipati osmolskae; Nank: Nankangia jiangxiensis; Anom: Anomalipes zhaoi; Neme: Nemegtia barsboldi; Anzu: Anzu wyliei; Giga: Gigantoraptor erlianensis.

Yilun Yu, Kebai Wang, Shuqing Chen, Corwin Sullivan, Shuo Wang, Peiye Wang and Xing Xu. 2018. A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Comments on Size Variation Among Oviraptorosaurs. Scientific Reports. volume 8, Article number: 5030. DOI: 10.1038/s41598-018-23252-2 

 researchgate.net/publication/323961130_A_new_caenagnathid_dinosaur_from_the_Upper_Cretaceous_Wangshi_Group_of_Shandong_China

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

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شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

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كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.

[Paleontology • 2018] Bayannurosaurus perfectus • A Large-sized Basal Ankylopollexian from East Asia, Shedding Light on early Biogeographic History of Iguanodontia ---ScRaBBlE

Bayannurosaurus perfectus 

 Xu, Tan, Gao, Bao, Yin, Guo, Wang, Tan, Zhang & Xing, 2018

  DOI:  10.1016/j.scib.2018.03.016 

Abstract

A presumably mostly quadrupedal ankylopollexian iguanodontian, Bayannurosaurus perfectus gen. et sp. nov., is reported here, and is represented by an excellently well-preserved skeleton from the Lower Cretaceous Bayingebi Formation of Inner Mongolia, China. The diagnosis of the taxon includes several autapomorphies, notably a dorsally directed, strap-like posterodorsal process of the jugal and a horizontally oriented preacetabular process of the ilium. The nearly complete caudal series retains eight posterior-most caudals with procoelous, trapezoidal centra in dorsal view, and the last three caudals are fully fused. The discovery of B. perfectus opens a critical new window on the early evolution and intercontinental dispersal of Iguanodontia. The skeleton displays a transitional morphology between non-hadrosauriform ankylopollexians and Hadrosauriformes. A phylogenetic analysis indicates that Bayannurosaurus is positioned higher on the tree than Hypselospinus, but below Ouranosaurus just outside of Hadrosauriformes. The tree topology of Iguanodontia with temporal and spatial constraints reveals a possible biogeographic scenario supported by the statistical dispersal-vicariance analysis: around the J/K boundary, non-hadrosauriform ankylopollexians experienced multiple dispersal events from Europe to Asia, accompanying the coeval fall of the global sea level.

Keywords: Non-hadrosauriform ankylopollexian; Lower Cretaceous; Bayingebi Formation; Inner Mongolia; Intercontinental dispersal

Fig. 1. Locality and horizon of the holotype (IMMNH PV00001) of Bayannurosaurus perfectus gen. et sp. nov.. (a) Fossil site at the quarry indicated by a white arrow. (b) Chulumiao locality in southwestern Inner Mongolia (China), where the holotype of B. perfectus was collected. (c) Stratigraphic distribution of B. perfectus within the upper Bayingebi Formation. (d) Holotype of B. perfectus originally exposed at the quarry.

Systematic palaeontology

Ornithischia Seeley, 1887

Ornithopoda Marsh, 1881

Iguanodontia Dollo, 1888 sensu Sereno, 1998

Ankylopollexia Sereno, 1986 sensu Sereno, 1998

Bayannurosaurus perfectus gen. et sp. nov.

 Etymology: ‘Bayannur’, the name of the general geographic area where the fossil was retrieved; ‘saurus’, based on the Greek word ‘sauros’ for lizard. ‘perfectus’, in reference to the perfect preservation of the skeleton designated as the holotypic specimen.

Holotype: IMMNH PV00001 (housed at the Inner Mongolia Museum of Natural History, Hohhot, Inner Mongolia), a nearly complete, semi-articulated skeleton, including the entire caudal series.

 Locality and horizon: Chulumiao, ∼7 km north of Chaogewenduer Town, Urad Rear Banner, Bayannur, Inner Mongolia (Fig. 1b); middle part of the upper half of the Bayingebi Formation (Fig. 1c); Early Cretaceous (early Aptian).

Xing Xu, Qingwei Tan, Yilong Gao, Zhiqiang Bao, Zhigang Yin, Bin Guo, Junyou Wang, Lin Tan, Yuguang Zhang and Hai Xing. 2018. A Large-sized Basal Ankylopollexian from East Asia, Shedding Light on early Biogeographic History of Iguanodontia. Science Bulletin.  63(9); 556-563. DOI:  10.1016/j.scib.2018.03.016

Researchgate.net/publication/324488089_A_large-sized_basal_ankylopollexian_from_East_Asia 

Iguanodontia is among the first-discovered dinosaurian groups in the world, and has become one of the best studied ornithischian clades, which refers to its taxonomic, phylogenetic and ontogenetic significance for dinosaurian evolution. Ankylopollexia represents a relatively derived iguanodontian clade that displays large body size and many highly modified features for more efficient herbivory and locomotion. Although relevant studies on European and North American material enhanced our knowledge of the early evolution of Ankylopollexia, the extreme scarcity of non-hadrosauriform ankylopollexians in Asia had long hindered attempts to comprehensively elucidate evolutionary history and related issues of the global iguanodontians. The research team led by Dr. Xing Xu names and describes a new non-hadrosauriform ankylopollexian Bayannurosaurus perfectus represented by an excellently wellpreserved skeleton, from the Lower Cretaceous Bayingebi Formation, western Inner Mongolia, China. The discovery of this new taxon is very helpful in understanding the origin and early evolution of Hadrosauriformes, optimizing the phylogenetic framework of non-hadrosaurid iguanodontians and clarifying the global biogeographic history of Iguanodontia around the Jurassic–Cretaceous boundary (see the article by Xing Xu et al. on page 556).

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روابط التحميل والمشاهدة، الروابط المباشرة للتحميل

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شاهد هذا الفيديو القصير لطريقة التحميل البسيطة

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كيف تحصل على مدونة جاهزة بآلاف المواضيع والمشاركات من هنا

شاهد قناة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على اليوتيوب لمزيد من الشرح من هنا

رابط مدونة منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات في أي وقت حــــتى لو تم حذفها من هنا
شاهد صفحة منتدى مدونات بلوجر جاهزة بألاف المواضيع والمشاركات على الفيس بوك لمزيد من الشرح من هنا

تعرف على ترتيب مواضيع منتدى مدونات بلوجر جاهزة بآلاف المواضيع والمشاركات (حتى لا تختلط عليك الامور) من هنا

ملاحظة هامة: كل عمليات تنزيل، رفع، وتعديل المواضيع الجاهزة تتم بطريقة آلية، ونعتذر عن اي موضوع مخالف او مخل بالحياء مرفوع بالمدونات الجاهزة بآلاف المواضيع والمشاركات، ولكم ان تقوموا بحذف هذه المواضيع والمشاركات والطريقة بسيطة وسهلة. ــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــــسلامـ.